217 resultados para Johtajan ominaisuudet
Resumo:
The previous academic research on Finnish peacekeeping has clarified the operative and historical aspects of Finnish peacekeeping lacking the view of an individual who does the actual peacekeeping work. This research is based on the underlying theoretical assumption of human beings possessing different kinds of talents and intelligences creating a holistic entity. In this broad perspective spirituality was explored as an umbrella concept, as a holistic ability or talent, that can be explored as the deepest aspect of defining what it means to be human. The theoretical framework incorporated the concept of an intelligence, which is defined in Gardner's theory of multiple intelligences as the ability to solve problems, or to create products, that are valued within one or more cultural settings (Gardner, 1993, x). The viability of this theory was studied in the sample of Finnish peacekeepers. Spirituality in the theoretical and conceptual horizon was viewed as an extension of Gardner's theory of intelligences as one potential Gardnerian intelligence candidate. In addition to Gardner's theory, spirituality was explored as sensitivity which includes capacities such as sensing awareness, sensing mystery and sensing value (Hay, 1998). Also the practical aspects of spirituality were taken in account as shown in our everyday lives giving us the direction and influencing our social responsibilities and concerns (Bradford, 1995). Spirituality was explored also involving the element of the peacekeepers' community, the element of personal moral orientations and in the domain of religion and coping. The purpose of this research aimed in two dimensions. First, the aim was to outline the intelligence profile and the spiritual sensitivity profile of peacekeepers. Second, the aim was to understand qualitatively the nature of peacekeepers' spirituality The research interests were studied with different kinds of peacekeepers. Applying the mixed methods approach the research was conducted in two phases: first the former SFOR peacekeepers (N=6) were interviewed and the data was analysed. Inspired by the primary findings of these interviews, the data for the case-study of one peacekeeper was collected in co-operation with one former SFOR peacekeeper (N=1). In the second phase the data was collected from KFOR peacekeepers through the quantitative MI-Survey and the spiritual sensitivity survey (N=195). The quantitative method was used to outline the intelligence profile and the spiritual sensitivity profile of peacekeepers (N=195). In the mixed methods approach this method highlighted the general overview of intelligence traits and spiritual sensitivity of peacekeepers. In the mixed methods approach the qualitative method including interviews (N=6) and a case-study of one peacekeeper (N=1) increased subjective, qualitative information of spirituality of peacekeepers. The intelligence profile of peacekeepers highlighted the bodily-kinesthetic and interpersonal dimensions as the practical and social aspects of peacekeepers. Strong inter-item dependencies in the intrapersonal intelligence profile meant that peacekeepers possess a self-reflection and self-knowledge component and they reflect on deep psychological and philosophical issues. Regarding the spiritual sensitivity, peacekeepers found awareness-sensing, mystery-sensing, value-sensing and community-sensing important. The community-sensing emphasised a strong will to advance peace and to help people who are in need: things that are close to the heart of the peacekeepers. These results depicted practicality, being socially capable, and reflecting one's inner world as essential to peacekeepers. Moreover, spirituality as peacekeepers' moral endeavour became clearer because the sub-model of their community-sensing described morally charged destinations: advancing peace and helping people in need. In the qualitative findings peacekeepers articulated justice orientation and rule-following characterising the nature of peacekeepers' moral attitude and moral call (Kohlberg, 1969). An ethic of care (Gilligan, 1982) describes mainly female moral orientation, but the findings revealed that an ethic of care is also an important agent supporting strongly male peacekeepers in their aim to carry out qualitatively good peacekeeping work. The moral endeavour was voiced, when the role of religion in coping meant the assessment of the a way of life, a way of conduct, a way of being truthful to one's own values in confusing surroundings. The practical level of spiritual and religious contemplation was voiced as morally charged inner motivation to fulfil one's duties and at the same time to cope with various peacekeeping challenges. The results of different data sets were combined and interpreted as the moral endeavour, which characterises peacekeepers' spirituality. As the combining result, the perspective of peacekeepers' spirituality is considered moral or at least morally charged.
Resumo:
Vuonna 2006 Suomessa korvattiin 2331 potilasvahinkoa. Niistä maksettiin korvauksia yhteensä noin 27 miljoonaa euroa. Tässä tutkimuksessa tarkastellaan potilasvahinkoja lääketieteellisen toiminnan valossa. Tutkimuksessa pyritään vastaamaan kysymykseen: Millä eri tavoilla lääkärin toiminta voi epäonnistua potilasvahinkotapauksissa? Tutkimuksessa määriteltiin neljä kategoriaa, joiden avulla analysoitiin esimerkkitapauksia. Kategoriat käsittelivät hoitojärjestelmää, lääkärin toimintaan vaikuttavia normeja, toiminnan intentionaalisuutta sekä toiminnan suhdetta sen lopputuloksiin ja seurauksiin. Ensimmäisessä tutkimusluvussa määriteltiin kategoriat, joiden avulla lääketieteellistä toimintaa voi arvioida. Ensimmäiseksi analysoitiin hoitojärjestelmää, lääkärin toiminnan kontekstia. Hoitojärjestelmästä nostettiin esille potilas-lääkärisuhde, lääkärien välinen yhteistyö, potilaasta saatavat tiedot, toiminnan mahdollistavat resurssit sekä niin kutsuttu systeemivirhe. Hoitojärjestelmän toimiminen on hyvän hoidon ensimmäinen edellytys. Se vaatii eri terveydenhuollon ammattihenkilöiden saumatonta yhteistyötä.Toinen kategoria käsitteli lääkärin toimintaan vaikuttavia normeja. Georg Henrik von Wrightin teoksen Norm and Action normien jaottelua sovellettiin lääkärin toimintaan. Normien pääryhmään kuuluvat säännöt, määräykset ja ohjeet. Lisäksi voidaan määritellä pienempiä normiryhmiä, jotka sijaitsevat pääryhmien välimaastossa. Näihin pienempiin ryhmiin sijoittuvat tottumukset, moraaliset periaatteet sekä ideaalit säännöt. Kolmannessa kategoriassa käsiteltiin intentionaalisuutta osana toimintaa. Teorian päälähde oli G. E. M. Anscomben teos Intention. Toiminnan intentionaalisuutta analysoitiin Anscomben tunnetun, vesikaivon myrkytystä koskevan, esimerkin avulla. Siinä tekoon liittyvä sisäinen tieto on keskeinen intentiota määrittävä tekijä. Viimeisessä kategoriassa arvioitiin toimintaa von Wrightin mukaan. Esille nousi erityisesti kyky toiminnan perusedellytyksenä sekä toiminnan lopputulos ja seuraukset. Potilasvahingoissa on tärkeää arvioida myös potilasvahingon seurauksia, ei vain epäonnistuneen toimenpiteen lopputulosta. Toimintaa analysoitaessa oli myös otettava huomioon kenen näkökulmasta seurauksia arvioitiin, Potilas, lääkäri ja esimerkiksi osastonylilääkäri saattavat nähdä potilasvahingon seuraukset hyvin eri tavoilla.Toisessa tutkimusluvussa arvioitiin neljää Terveydenhuollon oikeusturvakeskuksen ja lääninhallitusten sosiaali- ja terveysosastojen kantelu- ja valvontaratkaisuista valittua esimerkkitapausta. Ensimmäinen käsitteli lääkärin toiminnan epäonnistumista synnytyksessä, toinen psyykkisistä ongelmista ja alkoholiriippuvuudesta kärsivän lääkärin ammatinharjoittamisoikeuksien rajoittamista, kolmas lääketutkimuksen johtajan vakavia virheitä ja neljäs Terveydenhuollon oikeusturvakeskuksen pysyvän asiantuntijan ristiriitaisia asiantuntijalausuntoja. Lääkärin toiminnan epäonnistuminen vaikuttaa helposti hoitojärjestelmään, jonka toimiminen perustuu työntekijöiden yhteistyöhön. Hoitovirheissä rikotaan yleensä lääketieteen sääntöjä. Potilasvahingon sattuessa lääkärin asema muuttuu normiauktoriteetista normisubjektiksi. Lääkäri saattaa toiminnallaan rikkoa myös moraalisia sääntöjä tai lääkärin ihannetta. Puhtaissa vahinkotapauksissa toiminnan lopputulos ei vastaa lääkärin intentiota, potilaan parantamista. Kaikissa potilasvahinkotapauksissa ei voida puhua vahingoista, vaan kyseessä voi olla myös tarkoituksellinen teko. Toiminnan lopputuloksella voi olla laajat seuraukset, jotka tulisi myös ottaa huomioon toimintaa arvioitaessa.
Resumo:
Plutarkhoksen ja Uuden testamentin ajan kristillisten kirjoittajien naiskuvaa ja näkemystä avioliitosta ei ole juurikaan vertailtu aiemmin. Molempien teokset on kirjoitettu suunnilleen samoihin aikoihin ja ne esittävät samanlaisen eettisen ideaalin avioliitosta ja puolisoiden rooleista siinä. Plutarkhoksen näkemys avioliitosta näkyy selvimmin hänen teoksessaan Neuvoja vastanaineille. Myös muista Plutarkhoksen teoksista löytyy mainintoja avioliitosta ja puolisoiden rooleista. Uuden testamentin maininnat avioliitosta jakautuvat useisiin eri teksteihin. Evankeliumit käsittelevät avioeroa. Paavali käsittelee avioliittoa yleensä ja seksuaalisuutta avioliitossa. Uuden testamentin myöhemmät kirjeet, varsinkin niiden huoneentaulut, ja apostolisten isien tekstit keskittyvät eniten puolisoiden rooleihin avioliitossa. Plutarkhoksen näkemys avioliitosta oli positiivinen. Tämä näkyy useissa hänen teoksistaan. Kristittyjen kirjoittajien näkemys avioliitosta ei ole niin yksiselitteisen positiivinen. Paavali ei perustele avioliittoa muuten kuin haureuden välttämisellä. Kuitenkin Efesolaiskirjeessä avioliitto käy metaforaksi Kristuksen ja kirkon suhteelle. Plutarkhos ja kristityt kirjoittajat eivät anna paljon ohjeita puolison valinnasta. Plutarkhoksen mukaan puolison valinnassa tulee katsoa hänen luonnettaan, ei varallisuutta tai ulkonäköä. Paavalin ja Ignatioksen mukaan on parempi, jos puoliso on myös kristitty. Kristityt antavat myös ohjeita seka-avioliittoon, jossa puoliso on ei-uskova. 1. Pietarin kirjeen mukaan vaimon tulee pysyä uskossaan, eikä pelätä miestään. Plutarkhoksen mukaan vaimon tulisi palvoa samoja jumalia kuin miehensäkin. Plutarkhos näyttää suhtautuvan naisten aloitteentekoon avioerossa kielteisemmin kuin miesten. Kristityt kirjoittajat suhtautuvat avioeroon kielteisesti, vaikka lieventävätkin Jeesuksen avioerokieltoa. Plutarkhos ei hyväksy miehen aviollista uskottomuutta, mutta antaa miehelle kuitenkin myönnytyksen, jos tämä on liian heikko kontrolloimaan itseään. Kristityt kirjoittajat suhtautuvat uskottomuuteen yksiselitteisen kielteisesti. Ohjeissa avioparille Plutarkhos korostaa avioliiton harmoniaa ja avioparin riitelemättömyyttä enemmän kuin kristityt kirjoittajat. Plutarkhoksen ja kristittyjen kirjoittajien ohjeet vaimolle ovat samanlaisia. Vaimon ihanteelliset ominaisuudet olivat samoja; naisen tuli olla siveellinen ja häveliäs sekä hiljainen. Sekä Plutarkhos että huoneentaulujen kirjoittajat olivat sitä mieltä, että naisen tulisi alistua miehelleen. Plutarkhos ei perustele alistumista mitenkään, kun taas kristityt kirjoittajat antavat aina jonkun perustelun. Yleisin perustelu on apologeettiset syyt. Myös miehelle annettavat ohjeet ovat samanlaiset. Miehen tulee hallita vaimoaan, mutta sekä Plutarkhos että kristityt kirjoittajat rajoittavat miehen hallintavaltaa. Mies toimii myös vaimonsa opettajana. Plutarkhos antaa enemmän ja yksityiskohtaisempia ohjeita seksistä kuin kristityt kirjoittajat. Kristityistä kirjoittajista ainoastaan Paavali paneutuu näihin kysymyksiin tarkemmin. Sekä Plutarkhoksen että Paavalin mukaan ihmisten piti suhtautua seksiin kunnioituksella ja välttää liiallista himoa.
Resumo:
Tutkin pro gradu -työssäni Yhdysvaltojen mustien kansalaisoikeusliikkeen käsittelyä Suomen kirkollisessa lehdistössä liikkeen keskeisimpinä toiminnan vuosina 1963–1968. Selvitän, miten kirkollinen lehdistö reagoi kansalaisoikeusliikkeeseen, ja mitkä olivat niitä asioita ja tapahtumia, joita lehdet toivat liikkeestä esille. Keskeistä työssäni on myös kansalaisoikeusliikkeen näkyvimmän johtajan Martin Luther King Juniorin lehdistössä saaman huomion analysointi. Olennaista tutkimuksen kannalta on selvittää, käsittelivätkö tutkimani lehdet kansalaisoikeusliikkeen toimintaa hengellisestä näkökulmasta vai pelkästään uutisina Yhdysvaltojen poliittisista tapahtumista. Käytännössä tähän liittyy myös kysymys siitä, koskettiko Yhdysvaltojen mustien ahdinko kirjoitusten mukaan suomalaista kristittyä. Tutkimukseni taustan kannalta tärkeää on selvittää mustien rotusorron historiaa Yhdysvalloissa sekä Kingin liikkeeseen tuomaa ajattelupohjaa. Tutkimukseni lähteinä käytän Kotimaan, Församlingsbladetin, Herättäjän, Sanan ja Etsijän numeroita vuosilta 1963–1968. Vuodelta 1967 lähdemateriaalia ei ole kuitenkaan löytynyt. Ajallisesti tutkimus sijoittuu kansalaisoikeusliikkeen organisoiman Birminghamin kampanjan alusta aina Kingin murhan aiheuttamaan kuohuntaan asti. Tutkimukseni on vahvasti sidottu kansalaisoikeusliikkeen keskeisiin tapahtumiin, joiden kautta esittelen liikkeen saamaa huomiota. Tämän takia en käykään, Etsijää lukuun ottamatta, järjestelmällisesti läpi kaikkia tutkimieni vuosien numeroita, vaan tutkittavat numerot ovat ilmestyneet aikoina, jolloin kansalaisoikeusliike toimi aktiivisesti ja näkyvästi. Tutkimukseni on kirkollisen lehdistön ja kansalaisoikeusliikkeen tutkimuksessa ainutlaatuinen. Liikkeen lehdistössä saamaa vastaanottoa ei ole Suomessa aiemmin tutkittu. Kansalaisoikeusliikettä koskeva tutkimus on keskittynyt lähinnä Kingin ajattelun kuvailuun. Merkityksellisen tutkimusaiheesta tekee myös se, että syvemmällä tasolla tutkimus kuvailee sitä, millaisena kirkon epäviralliset äänenkannattajat näkivät ajan, jolloin yhteiskunnan sosiaalisiin ongelmiin puuttuminen alettiin nähdä kristityn tehtävänä. Lehtien kansalaisoikeusliikkettä koskeva keskustelu onkin nähtävä osana muuta ajan kehitystä ja sen henkeä. Vuosikymmenen ajan henki ja aikakauden suuret muutokset näkyvät selkeästi tutkimukseni tuloksissa. Kaukaisesta tapahtumapaikastaan huolimatta Yhdysvaltojen mustien kansalaisoikeusliike sai varsin paljon huomiota tutkimissani lehdissä niin pakinoissa, uutisissa kuin raporteissa tapahtumapaikoiltakin. Liikkeen toimintaa ja erityisesti Kingiä liikkeen johtajana ihailtiin ja liikettä hyödynnettiin esimerkkinä siitä, miten suomalaisten kristittyjenkin tulisi suhtautua maailmaan. Useissa lehdissä koettiinkin tarpeelliseksi nostaa esille Suomen omia ”rotukysymyksiä”, joista keskeisimpänä nähtiin romanien asema. Ajan kehitykseen varauksellisesti suhtautuvaa Herättäjää lukuun ottamatta lehdet suhtautuivat hyvin suopeasti mustien kansalaisoikeusliikkeen tarjoamaan sosiaalisen herätykseen. Erityisesti Kotimaan päätoimittaja Simo Talvitie paneutui mustien rotusortoon ja sen tarjoamiin opetuksiin myös Suomen tasolla. Selityksenä lehtien mustien kansalaisoikeusliikkeen runsaalle käsittelylle ja niiden tavalle käsitellä liikettä nousee keskeisenä aikaa leimaava kansainvälistyminen. Median kehittyminen erityisesti television myötä ja matkustamisen helpottuminen mahdollistivat entistä enemmän muiden maiden tapahtumien seuraamisen ja niihin kantaa ottamisen. Aikaansa seuraavien kristittyjen tehtäväksi nähtiinkin maailman tapahtumista perillä oleminen ja myös oman yhteiskunnan epäkohtien esille tuominen. Nämä eivät tutkimukseni mukaan olleet vain radikaalin ylioppilasnuorison kiinnostuksen kohteita vaan vähitellen myös monen suomalaisen kristityn.
Resumo:
Constitutional politics in Russia, a conceptual history study of the constitutional rhetoric in the 20th century In April 2006 the Russian Constitution had its 100th anniversary. Following its late start, five constitutions have been adopted. The great number of constitutions is partly explained in my work by the fact that Russia s political system has changed many times, from one state system to another. From a monarchical state power, it changed first, with the October revolution, into the Russian Socialist Federal Soviet Republic, and after that, in 1924, into the Union of Soviet Socialist Republics. In 1991, the Russian Federation was established. The great number of constitutions can also be explained by the fact that in a one-party system, constitutional concepts became one of the most important instruments for introducing political programmes. When the political unity of the state was not only restricted by the Constitution, but also by the party ideology, the political debates concerning constitutional concepts were the key discussions for all the reformative pursuits of Soviet politics. It can be said that, in the Soviet Union, almost all political discourses dealt with constitutional concepts. In the context of restricted unity, the constitutional concepts were the most important tools to argue and create a basis for a new presentation and new political programmes. Thus, the basic feature of the Soviet political discourses has been a continuous competition regarding the constitutional concepts. By defining the constitutional concepts, a new, the political elites wanted especially to redefine, their own way, the traditional meanings of the October 1917 Revolution, and to differentiate them from those of the preceding period of power. From a methodological point of view, I argue that the Russian constitutional concepts make a conceptual historical approach very suitable, and change the focus on history. This approach studies history in contemporary contexts which follow after each other, and whose texts are the main research target. The picture of history is created through the interpretation of the original sources of contemporary contexts. Focusing on the dynamic and traditional characteristic of Russian constitutional concepts, I refer to a certain kind of value and the task of these concepts to justify and define the political and societal unity in every situation. This is done by combining the pursued future orientation of constitutional unity with the new acts of preservation of the traditional principles of the revolution. The different time layers of the constitutional concepts, the past, the present and the future, are the key aspects of storytelling in justifying the continuity and redefining the constitutional unity for the sake of reform. These aspects of constitutional concepts, in addition to all the other functions, have been the main elements of the argumentative structure of acting against opponents.
Resumo:
In What We Owe to Each Other, T.M. Scanlon formulated a new version of the ethical theory called contractualism. This theory took reasons considerations that count in favour of judgment-sensitive attitudes to be the fundamental normative notion. It then used normative reasons to first account for evaluative properties. For an object to be valuable, on this view, is for it to have properties that provide reasons to have favourable attitudes towards the bearer of value. Scanlon also used reasons to account for moral wrongness. His contractualism claims that an act is morally wrong if it is forbidden by any set of moral principles that no one could reasonably reject. My thesis consists of five previously published articles which attempt to clarify Scanlon s theory and to defend it against its critics. The first article defends the idea that normative reason-relations are fundamental against Joshua Gert. Gert argues that rationality is a more basic notion than reasons and that reasons can be analysed in terms of their rationally requiring and justifying dimensions. The second article explores the relationship between value and reasons. It defends Scanlon s view according to which reasons are the more basic than value against those who think that reasons are based on the evaluative realm. The last three articles defend Scanlon s views about moral wrongness. The first one of them discusses a classic objection to contractualist theories. This objection is that principles which no one could reasonably reject are redundant in accounting for wrongness. This is because we need a prior notion of wrongness to select those principles and because such principles are not required to make actions wrong or to provide reasons against wrong actions. The fourth article explores the distinctive reasons which contractualists claim there are for avoiding the wrong actions. The last article argues against the critics of contractualism who claim that contractualism has implausible normative consequences for situations related to the treatment of different-sized groups of people.
Resumo:
Economic and Monetary Union can be characterised as a complicated set of legislation and institutions governing monetary and fiscal responsibilities. The measures of fiscal responsibility are to be guided by the Stability and Growth Pact, which sets rules for fiscal policy and makes a discretionary fiscal policy virtually impossible. To analyse the effects of the fiscal and monetary policy mix, we modified the New Keynesian framework to allow for supply effects of fiscal policy. We show that defining a supply-side channel for fiscal policy using an endogenous output gap changes the stabilising properties of monetary policy rules. The stability conditions are affected by fiscal policy, so that the dichotomy between active (passive) monetary policy and passive (active) fiscal policy as stabilising regimes does not hold, and it is possible to have an active monetary - active fiscal policy regime consistent with dynamical stability of the economy. We show that, if we take supply-side effects into ac-count, we get more persistent inflation and output reactions. We also show that the dichotomy does not hold for a variety of different fiscal policy rules based on government debt and budget deficit, using the tax smoothing hypothesis and formulating the tax rules as difference equations. The debt rule with active monetary policy results in indeterminacy, while the deficit rule produces a determinate solution with active monetary policy, even with active fiscal policy. The combination of fiscal requirements in a rule results in cyclical responses to shocks. The amplitude of the cycle is larger with more weight on debt than on deficit. Combining optimised monetary policy with fiscal policy rules means that, under a discretionary monetary policy, the fiscal policy regime affects the size of the inflation bias. We also show that commitment to an optimal monetary policy not only corrects the inflation bias but also increases the persistence of output reactions. With fiscal policy rules based on the deficit we can retain the tax smoothing hypothesis also in a sticky price model.
Resumo:
This study explores strategic political steering after the New Public Management (NPM) reforms, with emphasis on the new role assigned to Government ministers in Finland. In the NPM model, politicians concentrate on broad, principal issues, while agencies have discretion within the limits set by politicians. In Finland, strategic steering was introduced with Management by Results (MBR), but the actual tools for strategic political steering have been the Government Programme, the Government Strategy Portfolio (GSP) and Frame Budgeting. This study addresses these tools as means of strategic steering conducted by the Cabinet and individual ministers within their respective ministries. The time frame of the study includes the two Lipponen Cabinets between 1995 and 2003. Interviews with fourteen ministers as well as with fourteen top officials were conducted. In addition, administrative reform documents and documents related to strategic steering tools were analysed. The empirical conclusions of the study can be summarised as follows: There were few signs of strategic political steering in the Lipponen Cabinets. Although the Government Programmes of both Cabinets introduced strategic thinking, the strategic guidelines set forth at the beginning of the Programme were not linked to the GSP or to Frame Budgeting. The GSP could be characterised as the collected strategic agendas of each ministry, while there was neither the will nor the courage among Cabinet members to prioritise the projects and to make selections. The Cabinet used Frame Budgeting mainly in the sense of spending limits, not in making strategic allocation decisions. As for the GSP at the departmental level, projects were suggested by top officials, and ministers only approved the suggested list. Frame Budgeting at the departmental level proved to be the most interesting strategic steering tool from ministers viewpoint: they actively participated in defining which issues would need extra financing. Because the chances for extra financing were minimal, ministers had an effect only on a marginal share of the budget. At the departmental level, the study shows that strategic plans were considered the domain of officials. As for strategies concerning specific substances, there was variation in the interest shown by the ministers. A few ministers emphasised the importance of strategic work and led strategy processes. In most cases, however, officials led the process while ministers offered comments on the drafts of strategy documents. The results of this study together with experiences reported in other countries and local politics show that political decision-makers have difficulty operating at the strategic level. The conclusion is that politicians do not have sufficient incentive to perform the strategic role implied by the NPM type of reforms. Overall, the empirical results of the study indicate the power of politics over management reforms.
Resumo:
The results presented in this thesis show that all females of a given population do not necessarily choose similar mating partners. Specifically, partner preferences of a fish, the sand goby (Pomatoschistus minutus), varied among individual females and depended on the social context at the time of choice. I also show that females assess multiple mate choice cues simultaneously; partner preferences were based more strongly on an interaction effect between different choice cues than on any individual cue. Furthermore, I found that preferred matings involved fitness benefits in the form of increased offspring success, but these benefits were not significantly affected by mate compatibility. Hence, mate choice for partner compatibility does not appear to be an important determinant of the observed variation in female mate preferences in this species. The context-dependency of female mating preferences revealed is relevant to how genetic variation in sexually selected traits might be maintained: as the mating success of a certain male type varies according to the choice context, directional sexual selection on male traits is shown to be less intense than generally thought making for a slower loss of genetic variation in these traits. Mating preferences of sand gobies were assessed by giving females a binary choice between males that differed in body size and/or other focus traits. These association preferences were found to be sexually motivated, repeatable and to correspond to actual mating decisions.
Resumo:
Vehicles affect the concentrations of ambient airborne particles through exhaust emissions, but particles are also formed in the mechanical processes in the tire-road interface, brakes, and engine. Particles deposited on or in the vicinity of the road may be re-entrained, or resuspended, into air through vehicle-induced turbulence and shearing stress of the tires. A commonly used term for these particles is road dust . The processes affecting road dust emissions are complex and currently not well known. Road dust has been acknowledged as a dominant source of PM10 especially during spring in the sub-arctic urban areas, e.g. in Scandinavia, Finland, North America and Japan. The high proportion of road dust in sub-arctic regions of the world has been linked to the snowy winter conditions that make it necessary to use traction control methods. Traction control methods include dispersion of traction sand, melting of ice with brine solutions, and equipping the tires with either metal studs (studded winter tires), snow chains, or special tire design (friction tires). Several of these methods enhance the formation of mineral particles from pavement wear and/or from traction sand that accumulate in the road environment during winter. When snow and ice melt and surfaces dry out, traffic-induced turbulence makes some of the particles airborne. A general aim of this study was to study processes and factors underlying and affecting the formation and emissions of road dust from paved road surfaces. Special emphasis was placed on studying particle formation and sources during tire road interaction, especially when different applications of traction control, namely traction sanding and/or winter tires were in use. Respirable particles with aerodynamic diameter below 10 micrometers (PM10) have been the main concern, but other size ranges and particle size distributions were also studied. The following specific research questions were addressed: i) How do traction sanding and physical properties of the traction sand aggregate affect formation of road dust? ii) How do studded tires affect the formation of road dust when compared with friction tires? iii) What are the composition and sources of airborne road dust in a road simulator and during a springtime road dust episode in Finland? iv) What is the size distribution of abrasion particles from tire-road interaction? The studies were conducted both in a road simulator and in field conditions. The test results from the road simulator showed that traction sanding increased road dust emissions, and that the effect became more dominant with increasing sand load. A high percentage of fine-grained anti-skid aggregate of overall grading increased the PM10 concentrations. Anti-skid aggregate with poor resistance to fragmentation resulted in higher PM levels compared with the other aggregates, and the effect became more significant with higher aggregate loads. Glaciofluvial aggregates tended to cause higher particle concentrations than crushed rocks with good fragmentation resistance. Comparison of tire types showed that studded tires result in higher formation of PM emissions compared with friction tires. The same trend between the tires was present in the tests with and without anti-skid aggregate. This finding applies to test conditions of the road simulator with negligible resuspension. Source and composition analysis showed that the particles in the road simulator were mainly minerals and originated from both traction sand and pavement aggregates. A clear contribution of particles from anti-skid aggregate to ambient PM and dust deposition was also observed in urban conditions. The road simulator results showed that the interaction between tires, anti-skid aggregate and road surface is important in dust production and the relative contributions of these sources depend on their properties. Traction sand grains are fragmented into smaller particles under the tires, but they also wear the pavement aggregate. Therefore particles from both aggregates are observed. The mass size distribution of traction sand and pavement wear particles was mainly coarse, but fine and submicron particles were also present.
Resumo:
Environmental variation is a fact of life for all the species on earth: for any population of any particular species, the local environmental conditions are liable to vary in both time and space. In today's world, anthropogenic activity is causing habitat loss and fragmentation for many species, which may profoundly alter the characteristics of environmental variation in remaining habitat. Previous research indicates that, as habitat is lost, the spatial configuration of remaining habitat will increasingly affect the dynamics by which populations are governed. Through the use of mathematical models, this thesis asks how environmental variation interacts with species properties to influence population dynamics, local adaptation, and dispersal evolution. More specifically, we couple continuous-time continuous-space stochastic population dynamic models to landscape models. We manipulate environmental variation via parameters such as mean patch size, patch density, and patch longevity. Among other findings, we show that a mixture of high and low quality habitat is commonly better for a population than uniformly mediocre habitat. This conclusion is justified by purely ecological arguments, yet the positive effects of landscape heterogeneity may be enhanced further by local adaptation, and by the evolution of short-ranged dispersal. The predicted evolutionary responses to environmental variation are complex, however, since they involve numerous conflicting factors. We discuss why the species that have high levels of local adaptation within their ranges may not be the same species that benefit from local adaptation during range expansion. We show how habitat loss can lead to either increased or decreased selection for dispersal depending on the type of habitat and the manner in which it is lost. To study the models, we develop a recent analytical method, Perturbation expansion, to enable the incorporation of environmental variation. Within this context, we use two methods to address evolutionary dynamics: Adaptive dynamics, which assumes mutations occur infrequently so that the ecological and evolutionary timescales can be separated, and via Genotype distributions, which assume mutations are more frequent. The two approaches generally lead to similar predictions yet, exceptionally, we show how the evolutionary response of dispersal behaviour to habitat turnover may qualitatively depend on the mutation rate.
Resumo:
Individuals face variable environmental conditions during their life. This may be due to migration, dispersion, environmental changes or, for example, annual variation in weather conditions. Genetic adaptation to a novel environment happens through natural selection. Phenotypic plasticity allows, however, a quick individual response to a new environment. Phenotypic plasticity may also be beneficial for individual if the environment is highly variable. For example, eggs are costly to produce. If the food conditions vary significantly between breeding seasons it is useful to be able to adjust the clutch and egg size according to the food abundance. In this thesis I use Ural owl vole system to study phenotypic plasticity and natural selection using a number of reproduction related traits. The Ural owl (Strix uralensis) is a long-lived and sedentary species. The reproduction and survival of the Ural owl, in fact their whole life, is tied to the dramatically fluctuating vole densities. Ural owls do not cause vole cycles but they have to adjust their behaviour to the rather predictable population fluctuations of these small mammals. Earlier work with this system has shown that Ural owl laying date and clutch size are plastic in relation to vole abundance. Further, individual laying date clutch size reaction norms have been shown to vary in the amount of plasticity. My work extends the knowledge of natural selection and phenotypic plasticity in traits related to reproduction. I show that egg size, timing of the onset of incubation and nest defense aggressiveness are plastic traits with fitness consequences for the Ural owl. Although egg size is in general thought to be a fixed characteristic of an individual, this highly heritable trait in the Ural owl is also remarkably plastic in relation to the changes in vole numbers, Ural owls are laying the largest eggs when their prey is most abundant. Timing of the onset of incubation is an individual-specific property and plastic in relation to clutch size. Timing of incubation is an important underlying cause for asynchronous hatching in birds. Asynchronous hatching is beneficial to offspring survival in Ural owl. Hence, timing of the onset of incubation may also be under natural selection. Ural owl females also adjust their nest defense aggressiveness according to the vole dynamics, being most aggressive in years when they produce the largest broods. Individual females show different levels of nest defense aggressiveness. Aggressiveness is positively correlated with the phenotypic plasticity of aggressiveness. As elevated nest defense aggressiveness is selected for, it may promote the plasticity of aggressive nest defense behaviour. All the studied traits are repeatable or heritable on individual level, and their expression is either directly or indirectly sensitive to changes in vole numbers. My work considers a number of important fitness-related traits showing phenotypic plasticity in all of them. Further, in two chapters I show that there is individual variation in the amount of plasticity exhibited. These findings on plasticity in reproduction related traits suggest that variable environments indeed promote plasticity.
Resumo:
Physicochemical characterization of freshwater samples from Finland, Sweden, the Netherlands, and Spain revealed that water hardness and pH decreased and the quantity and quality of humic substances changed considerably in this geographical series from south to north. Since the ambient water chemistry may affect the availability of chemicals, the total aqueous concentration of a chemical may be insufficient to predict the bioconcentration, subsequent biological response, and thus risk. In addition, organisms could be affected directly by water quality characteristics. In this context the main objective of this thesis was to investigate the bioavailability of selected ecotoxicologically relevant chemicals (cadmium, benzo(a)pyrene, and pyrene) in various European surface waters and to show the importance of certain water chemistry characteristics in interpreting the bioavailability and toxicity results. The bioavailability of cadmium to Daphnia magna was examined in very soft humic lake water. Humic substances as natural ligands decreased the free and bioavailable proportion of cadmium in soft lake water. As a consequence the uptake rate and the acute toxicity decreased compared with the humic-free reference. When the hardness of humic lake water was artificially elevated, the acute toxicity of cadmium decreased, although the proportion of free cadmium increased. The decreased bioavailability of cadmium in hard water was a result of effective competition for uptake by the hardness cations, especially calcium ions. The protective role of humic substances and water hardness against cadmium toxicity was also observed in Lumbriculus variegatus, although D. magna was more sensitive to cadmium. The bioavailability of two polycyclic aromatic hydrocarbons (PAHs), pyrene and benzo(a)pyrene, was studied in European surface waters of varying water chemistry. Humic substances acted as complexing ligands with both PAHs, but the bioavailability of the more lipophilic benzo(a)pyrene to D. magna was affected more by humic substances than that of pyrene. In addition, not only the quantity of humic substances, but also their quality affected the bioavailability of benzo(a)pyrene. Nevertheless, the humic substances played a protective role in the photo-enhanced toxicity of pyrene under UV-B radiation. Water hardness had no effect on pyrene toxicity. Results indicate that the typical physicochemical characteristics of boreal freshwaters should be considered carefully in local and regional risk assessment of chemicals concerning the Fennoscandian region.
Resumo:
Ongoing habitat loss and fragmentation threaten much of the biodiversity that we know today. As such, conservation efforts are required if we want to protect biodiversity. Conservation budgets are typically tight, making the cost-effective selection of protected areas difficult. Therefore, reserve design methods have been developed to identify sets of sites, that together represent the species of conservation interest in a cost-effective manner. To be able to select reserve networks, data on species distributions is needed. Such data is often incomplete, but species habitat distribution models (SHDMs) can be used to link the occurrence of the species at the surveyed sites to the environmental conditions at these locations (e.g. climatic, vegetation and soil conditions). The probability of the species occurring at unvisited location is next predicted by the model, based on the environmental conditions of those sites. The spatial configuration of reserve networks is important, because habitat loss around reserves can influence the persistence of species inside the network. Since species differ in their requirements for network configuration, the spatial cohesion of networks needs to be species-specific. A way to account for species-specific requirements is to use spatial variables in SHDMs. Spatial SHDMs allow the evaluation of the effect of reserve network configuration on the probability of occurrence of the species inside the network. Even though reserves are important for conservation, they are not the only option available to conservation planners. To enhance or maintain habitat quality, restoration or maintenance measures are sometimes required. As a result, the number of conservation options per site increases. Currently available reserve selection tools do however not offer the ability to handle multiple, alternative options per site. This thesis extends the existing methodology for reserve design, by offering methods to identify cost-effective conservation planning solutions when multiple, alternative conservation options are available per site. Although restoration and maintenance measures are beneficial to certain species, they can be harmful to other species with different requirements. This introduces trade-offs between species when identifying which conservation action is best applied to which site. The thesis describes how the strength of such trade-offs can be identified, which is useful for assessing consequences of conservation decisions regarding species priorities and budget. Furthermore, the results of the thesis indicate that spatial SHDMs can be successfully used to account for species-specific requirements for spatial cohesion - in the reserve selection (single-option) context as well as in the multi-option context. Accounting for the spatial requirements of multiple species and allowing for several conservation options is however complicated, due to trade-offs in species requirements. It is also shown that spatial SHDMs can be successfully used for gaining information on factors that drive a species spatial distribution. Such information is valuable to conservation planning, as better knowledge on species requirements facilitates the design of networks for species persistence. This methods and results described in this thesis aim to improve species probabilities of persistence, by taking better account of species habitat and spatial requirements. Many real-world conservation planning problems are characterised by a variety of conservation options related to protection, restoration and maintenance of habitat. Planning tools therefore need to be able to incorporate multiple conservation options per site, in order to continue the search for cost-effective conservation planning solutions. Simultaneously, the spatial requirements of species need to be considered. The methods described in this thesis offer a starting point for combining these two relevant aspects of conservation planning.
Resumo:
Wild salmon stocks in the northern Baltic rivers became endangered in the second half of the 20th century, mainly due to recruitment overfishing. As a result, supplementary stocking was widely practised, and supplementation of the Tornionjoki salmon stock took place over a 25 year period until 2002. The stock has been closely monitored by electrofishing, smolt trapping, mark-recapture studies, catch samples and catch surveys. Background information on hatchery-reared stocked juveniles was also collected for this study. Bayesian statistics was applied to the data as this method offers the possibility of bringing prior information into the analysis and an advanced ability for incorporating uncertainty, and also provides probabilities for a multitude of hypotheses. Substantial divergences between reared and wild Tornionjoki salmon were identified in both demographic and phenological characteristics. The divergences tended to be larger the longer the duration spent in hatchery and the more favourable the hatchery conditions were for fast growth. Differences in environment likely induced most of the divergences, but selection of brood fish might have resulted in genotypic divergence in maturation age of reared salmon. Survival of stocked 1-year old juveniles to smolt varied from about 10% to about 25%. Stocking on the lower reach of the river seemed to decrease survival, and the negative effect of stocking volume on survival raises the concern of possible similar effects on the extant wild population. Post-smolt survival of wild Tornionjoki smolts was on average two times higher than that of smolts stocked as parr and 2.5 times higher than that of stocked smolts. Smolts of different groups showed synchronous variation and similar long-term survival trends. Both groups of reared salmon were more vulnerable to offshore driftnet and coastal trapnet fishing than wild salmon. Average survival from smolt to spawners of wild salmon was 2.8 times higher than that of salmon stocked as parr and 3.3 times higher than that of salmon stocked as smolts. Wild salmon and salmon stocked as parr were found to have similar lifetime survival rates, while stocked smolts have a lifetime survival rate over 4 times higher than the two other groups. If eggs are collected from the wild brood fish, stocking parr would therefore not be a sensible option. Stocking smolts instead would create a net benefit in terms of the number of spawners, but this strategy has serious drawbacks and risks associated with the larger phenotypic and demographic divergences from wild salmon. Supplementation was shown not to be the key factor behind the recovery of the Tornionjoki and other northern Baltic salmon stocks. Instead, a combination of restrictions in the sea fishery and simultaneous occurrence of favourable natural conditions for survival were the main reasons for the revival in the 1990 s. This study questions the effectiveness of supplementation as a conservation management tool. The benefits of supplementation seem at best limited. Relatively high occurrences of reared fish in catches may generate false optimism concerning the effects of supplementation. Supplementation may lead to genetic risks due to problems in brood fish collection and artificial rearing with relaxed natural selection and domestication. Appropriate management of fisheries is the main alternative to supplementation, without which all other efforts for long-term maintenance of a healthy fish resource fail.