A history of the British freshwater Algæ, including descriptions of the Desmideæ and Diatomaceæ ... By Arthur Hill Hassall ...

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Paleoenvironmental changes during the late Miocene (Messinian)–Pliocene transition (Bajo Segura Basin, southeastern Spain): sedimentological and ichnological evidence

A detailed sedimentological and paleontological analysis of the uppermost Miocene (Messinian)–Pliocene boundary at the northern border of the Bajo Segura Basin, southeastern Spain, was carried out in order to describe the evolution of the regional paleocoastline during the Pliocene reflooding of the Mediterranean immediately after the sea-level fall related to the Messinian Salinity Crisis. Multiple trace fossils typical of firm- and hardgrounds were recognized, allowing identification of Glossifungites (two different types), Entobia, and Gnathichnus ichnofacies. Trace-fossil analysis showed that lithology and media consistency exerted considerable control on the development of the different ichnocoenoses and that there was a clear decrease in hydrodynamic energy from a coastal to a shallow-water shelf environment related to progressive sea-level rise. Ichnological and sedimentological data provide evidence that the definitive flooding of the Mediterranean was rapid and synchronous throughout the northern margin of the Bajo Segura Basin. The following model for the Pliocene transgression in the study area is therefore proposed: (1) the marine ingression penetrated along the incised paleovalleys carved as a consequence of the fall in sea level, where the first two Pliocene systems were deposited (P0–P1); (2) during the maximum flooding surface of the transgression, the sea overflowed the margins of the paleovalleys and extended throughout the entire northern margin of the basin; and (3) the third Pliocene system was deposited, forming the lower part of a highstand systems tract (P2).

Integrated taphonomy in an open-marine platform: The Lower Cretaceous of Sierra Helada (Betic Cordillera, SE Spain)

A detailed taphonomic analysis was carried out on the lower Albian deposits of the Sierra Helada section (Alicante, Betic Cordillera, southeastern Spain). Ten taphonomic characters were studied and ten skeletal concentrations were defined on the basis of taphonomic features and the dominant taxa. Cluster analysis was performed on the dataset represented by the abundance of the taphonomic characters in each skeletal concentration. This enabled the definition of four different taphonomic categories: 1) skeletal concentrations characterized by the presence of fossils preserved in life position, 2) skeletal concentrations showing very little physical reworking, 3) skeletal concentrations related to high-energy background conditions, and 4) skeletal concentrations produced by medium- to high-energy events. Four taphofacies were defined on the basis of the main sedimentological features and the most representative skeletal concentrations. Taphofacies A represents the low energy outer platform, rich in skeletal concentrations with echinoids in life position and only slightly reworked. The second taphofacies (taphofacies B) is very rich in reworked echinoid tests and calcarenitic beds and records the transition to shallower areas, while taphofacies C shows abundant thick-bedded calcarenites and skeletal concentrations produced by sediment transport and rapid deposition. Finally, cross-bedded grainstone beds, which are rich in fine-grained fragmented, locally reoriented bioclasts (taphofacies D), record the existence of shifting sandy dunes in the shallow inner part of the platform.

Pollen abundances, and temperature and precipitation reconstruction for the Eocene to Oligocene transition in northern high latitudes

A profound global climate shift took place at the Eocene-Oligocene transition (~33.5 million years ago) when Cretaceous/early Palaeogene greenhouse conditions gave way to icehouse conditions (Zachos et al., 2001, doi:10.1126/science.1059412; Coxall et al., 2005, doi:10.1038/nature03135; Lear et al., 2008, doi:10.1130/G24584A.1). During this interval, changes in the Earth's orbit and a long-term drop in atmospheric carbon dioxide concentrations (Pagani et al., 2005, doi:10.1126/science.1110063; Pearson and Palmer, 2000, doi:10.1038/35021000; DeConto and Pollard, 2003, doi:10.1038/nature01290) resulted in both the growth of Antarctic ice sheets to approximately their modern size (Coxall et al., 2005, doi:10.1038/nature03135; Lear et al., 2008, doi:10.1130/G24584A.1) and the appearance of Northern Hemisphere glacial ice (Eldrett et al., 2007, doi:10.1038/nature05591; Moran et al., 2006, doi:10.1038/nature04800). However, palaeoclimatic studies of this interval are contradictory: although some analyses indicate no major climatic changes (Kohn et al., 2004, doi:10.1130/G20442.1; Grimes et al., 2005, doi:10.1130/G21019.1), others imply cooler temperatures (Zanazzi et al., 2007, doi:10.1038/nature05551), increased seasonality (Ivany et al., 2000, doi:10.1038/35038044; Terry, 2001, doi:10.1016/S0031-0182(00)00248-0) and/or aridity (Ivany et al., 2000, doi:10.1038/35038044; Terry, 2001, doi:10.1016/S0031-0182(00)00248-0; Sheldon et al., 2002, doi:10.1086/342865; Dupont-Nivet et al., 2007, doi:10.1038/nature05516). Climatic conditions in high northern latitudes over this interval are particularly poorly known. Here we present northern high-latitude terrestrial climate estimates for the Eocene to Oligocene interval, based on bioclimatic analysis of terrestrially derived spore and pollen assemblages preserved in marine sediments from the Norwegian-Greenland Sea. Our data indicate a cooling of ~5 °C in cold-month (winter) mean temperatures to 0-2 °C, and a concomitant increased seasonality before the Oi-1 glaciation event. These data indicate that a cooling component is indeed incorporated in the d18O isotope shift across the Eocene-Oligocene transition. However, the relatively warm summer temperatures at that time mean that continental ice on East Greenland was probably restricted to alpine outlet glaciers.

Tropical Atlantic SST during the mid-Pleistocene transition (MPT) of ODP Hole 175-1077B

We compare a new mid-Pleistocene sea surface temperature (SST) record from the eastern tropical Atlantic to changes in continental ice volume, orbital insolation, Atlantic deepwater ventilation, and Southern Ocean front positions to resolve forcing mechanisms of tropical Atlantic SST during the mid-Pleistocene transition (MPT). At the onset of the MPT, a strong tropical cooling occurred. The change from a obliquity- to a eccentricity-dominated cyclicity in the tropical SST took place at about 650 kyr BP. In orbital cycles, tropical SST changes significantly preceded continental ice-volume changes but were in phase with movements of Southern Ocean fronts. After the onset of large-amplitude 100-kyr variations, additional late glacial warming in the eastern tropical Atlantic was caused by enhanced return flow of warm waters from the western Atlantic driven by strong trade winds. Pronounced 80-kyr variations in tropical SST occurred during the MPT, in phase with and likely directly forced by transitional continental ice-volume variations. During the MPT, a prominent anomalous long-term tropical warming occurred, likely generated by extremely northward displaced Southern Ocean fronts. While the overall pattern of global climate variability during the MPT was determined by changes in mean state and frequency of continental ice volume variations, tropical Atlantic SST variations were primarily driven by early changes in Subantarctic sea-ice extent and coupled Southern Ocean frontal positions.

(Table 1) Age and depth of reflectors of the 3.5-kHz and airgun seismic record and their corresponding depth of ash layers, carbonate peaks, and transition zones from DSDP Hole 83-504B

DSDP cores from areas of low (Site 505) and high heat flow (Site 504 B) near the Costa Rica Rift, together with seismic profiles from the Panama Basin, have been studied to determine the relationship between: (1) carbonate content and physical and acoustic properties; and (2) carbonate content, carbonate diagenesis and acoustic stratigraphy. Except for ash and chert layers, bulk density correlates strongly and linearly with carbonate content. Velocity is uniform downcore and only small variations at a small scale are measured. Thus an abrupt change in carbonate content will cause abrupt changes in acoustic impedance and should cause reflectors that can be detected acoustically. A comparison of seismic profiler reflection records with physical properties, carbonate content and reflection coefficients indicates that the main reflectors can be identified with ash layers, diagenetic boundaries, and carbonate content variations. Diagenesis of carbonate sediments is present at Site 504B in a 260 m-thick ooze-chalk-limestone/chert sequence. These diagenetic sequences occur in areas of higher heat flow (200 mW/m**2). Seismic profiler records can be used to map the extent and depth of these diagenetic boundaries.

Radiocarbon data, oxygen isotopic composition in foraminifera, freshwater diatoms data, and salinity estimations from ODP Holes 167-1019C and 167-1019D

Massive discharges of freshwater from the glacial lake Missoula to the northeast Pacific Ocean are thought to have sculpted the Channeled Scablands of eastern Washington and debouched via the Columbia River near 46°N. The dynamics and timing of these events and their impact on northeast Pacific circulation remain uncertain. Here we date marine records of anomalous freshwater inputs to the ocean based on freshwater diatoms, oxygen isotopes in foraminifera, and radiocarbon data. Low-salinity plumes from the Columbia River reduced sea-surface salinities by as much as 6 psu (practical salinity units) more than 400 km away between 16 and 31 cal (calendar) ka B.P. Anomalously high abundances of freshwater diatoms in marine sediments from the region precede generally accepted dates for the existence of glacial Lake Missoula, implying that large flooding or freshwater routing events were common during the advance of the Cordilleran Ice Sheet and that such events require multiple sources.

Marine ice-rafted debris records and age determinations along the northern Eurasian margin

Ice-rafted debris (IRD) (>2 mm), input in eight sediment cores along the Eurasian continental margin (Arctic Ocean), have been studied over the last two glacial/interglacial cycles. Together with the revised chronologies and new micropaleontological data of two cores from the northern Barents Sea (PS2138) and northeastern Kara Sea (PS2741) spanning Marine Isotope Stages (MIS) 6 to 1, the IRD data give new insights into the glacial history of northern Eurasian ice-sheets over the last 150 ka. The chronologies of the cores are based on stable isotope records, AMS 14C datings, paleomagnetic and biostratigraphic data. Extensive episodes of northern Barents Sea ice-sheet growth, probably to the shelf edge, occurred during the late Weichselian (MIS 2) and the Saalian (MIS 6). Major IRD discharge at the MIS 4/3-transition hints to another severe glaciation, probably onto the outer shelf, during MIS 4. IRD-based instabilities of the marine-based ice margin along the northern Barents Sea between MIS 4 and 2 are similar in timing with North Atlantic Heinrich events and Nordic Seas IRD events, suggesting similar atmospheric cooling over a broad region or linkage of ice-sheet fluctuations through small sea-level events. In the relatively low-precipitation areas of eastern Eurasia, IRD peak values during Termination II and MIS 4/3-transition suggest a Kara Sea ice-sheet advance onto the outer shelf, probably to the shelf edge, during glacial MIS 6 and 4. This suggests that during the initial cooling following the interglacials MIS 5, and possibly MIS 7, the combined effect of sustained inflow of Atlantic water into the Arctic Ocean and penetration of moisture-bearing cyclones into easterly direction supported major ice build-up during Saalian (MIS 6) and Mid-Weichselian (MIS 4) glaciation. IRD peak values in MIS 5 indicate at least two advances of the Severnaya Semlya ice-sheet to the coast line during the Early Weichselian. In contrast, a distinct Kara Sea ice advance during the Late Weichselian (MIS 2) is not documented by the IRD records along the northeastern Kara Sea margin.

Changes in the vegetation and trade winds in equatorial Northwest Africa 140,000-70,000 yr B.P., as deduced from two marine pollen records

The deep-sea cores M 16415-2 and M 16416-2 at about 9°N off Sierra Leone were analysed palynologically for the time interval 140,000-70,000 yr B.P. Results were presented in absolute (pollen concentration and pollen influx) and relative diagrams (pollen percentage). In a previous study it was evidenced that in northwest Africa pollen is mainly transported to the Atlantic by wind, so that the efficiency of aeolian pollen transport (pollen flux) could be used to evaluate changes in the intensity of the northeast trade winds. The glacial episodes (represented by the oxygen isotope stages 6 and 4) are characterized by strong northeast trade winds, whereas the last interglacial (stage 5) is characterized by weak trade winds. The pollen influx diagram shows that the intensity of the trade winds increased slightly during the relatively cool intervals of stage 5 (viz. 5.4 and 5.2). Tropical forest had maximally expanded around 124,000 yr B.P. (stage 5.5), around 98,000 yr B.P. (transition of stage 5.3 to 5.2), and around 70,000 yr B.P. (first part of stage 4): an increasing delay of the response of tropical forest to global intervals with maximum temperature is apparent during the last interglacial. As tropical forests need continuous humidity, the record of tropical forest monitors changes in climatic humidity south of the Sahara. During the last interglacial, the southern boundary of the Sahara shifted only little: expansions and contractions of the tropical forest area are correlated with contra-oscillations of the grass-dominated savanna zone. Great latitudinal shifts of the desert savanna boundary, on the contrary, occurred during the penultimate glacial interglacial transition (around 128,000 yr B.P.) to the north, and during the last interglacial-glacial transition (around 65,000 yr B.P.) to the south.

(Table S1) Age determination of sediment cores of the Gulf of Alaska

Oxygen isotope data from planktonic and benthic foraminifera, on a high-resolution age model (44 14C dates spanning 17,400 years), document deglacial environmental change on the southeast Alaska margin (59°33.32'N, 144°9.21'W, 682 m water depth). Surface freshening (i.e., d18O reduction of 0.8 per mil) began at 16,650 ± 170 cal years B.P. during an interval of ice proximal sedimentation, likely due to freshwater input from melting glaciers. A sharp transition to laminated hemipelagic sediments constrains retreat of regional outlet glaciers onto land circa 14,790 ± 380 cal years B.P. Abrupt warming and/or freshening of the surface ocean (i.e., additional d18O reduction of 0.9 per mil) coincides with the Bølling Interstade of northern Europe and Greenland. Cooling and/or higher salinities returned during the Allerød interval, coincident with the Antarctic Cold Reversal, and continue until 11,740 ± 200 cal years B.P., when onset of warming coincides with the end of the Younger Dryas. An abrupt 1 per mil reduction in benthic d18O at 14,250 ± 290 cal years B.P. likely reflects a decrease in bottom water salinity driven by deep mixing of glacial meltwater, a regional megaflood event, or brine formation associated with sea ice. Two laminated opal-rich intervals record discrete episodes of high productivity during the last deglaciation. These events, precisely dated here at 14,790 ± 380 to 12,990 ± 190 cal years B.P. and 11,160 ± 130 to 10,750 ± 220 cal years B.P., likely correlate to similar features observed elsewhere on the margins of the North Pacific and are coeval with episodes of rapid sea level rise. Remobilization of iron from newly inundated continental shelves may have helped to fuel these episodes of elevated primary productivity and sedimentary anoxia.

Organic matter deposition along the Kara and Laptev Seas continental margin (eastern Arctic Ocean) during last deglaciation and Holocene

Organic petrologic (maceral analysis) and bulk organic-geochemical studies were performed on five sediment cores from the Eurasian continental margin to reconstruct the environmental changes during the last not, vert, similar13 000 yr. The core stratigraphy is based on AMS-14C dating, and correlation by magnetic susceptibility and lithostratigraphic characteristics. Variations in terrigenous, freshwater, and marine organic matter deposition document paleoceanographic and paleoclimatic changes during the transition from the last deglaciation to the Holocene. Glacigenic diamictons deposited in the St. Anna Trough (northern Kara Sea) during the Last Glacial Maximum (LGM) are characterized by reworked terrigenous organic matter. In contrast, the Laptev Sea shelf was not covered by an ice-sheet, but was exposed by the lowered sea level. Increased deposition of marine organic matter (MOM) during deglaciation indicates enhanced surface-water productivity, possibly related to influence of Atlantic waters. The occurrence of freshwater alginite gives evidence for river discharge to the Kara and Laptev Seas after the LGM. At the eastern Laptev Sea slope, the first influence of Atlantic water masses is indicated by an increase in the contents of MOM and dinoflagellate cysts, with Operculodinium centrocarpum prior to not, vert, similar10 000 yr BP. High sedimentation rates in the Kara and the Laptev Seas with the adjacent slope at the beginning of the Holocene are presumably related to increased freshwater and sediment discharge from the Siberian rivers. Evidence for elevated Holocene freshwater discharge to the Laptev Sea has been found between not, vert, similar9.8 and 9 kyr BP, at not, vert, similar5 kyr BP and at not, vert, similar2.5 kyr BP. In the Kara Sea, an increased freshwater signal is obvious at not, vert, similar8.5 kyr BP and at not, vert, similar5 kyr BP. Higher portions of MOM were accumulated in the St. Anna Trough and at the Eurasian continental margin at several intervals during the Holocene. Increased primary productivity during these intervals is explained by seasonally ice-free conditions possibly associated with increased inflow of Atlantic waters.

Response and function of cutaneous mucosal and serum antibodies in barramundi (Lates calcarifer) acclimated in seawater and freshwater

Mucosal and serum antibody responses were studied in sibling barramundi (Lates calcarifer) acclimated in either seawater or freshwater following vaccination by intraperitoneal injection or direct immersion in an inactivated Streptococcus iniae vaccine. As expected, route of vaccination had a marked effect on immune response, with direct immersion resulting in low serum antibody levels against S. iniae by ELISA detected 21 days post vaccination at 26 degrees C, whilst a significant response was detected in mucus. A strong specific antibody response was detected in both mucus and serum 21 days following intraperitoneal injection. Fish acclimated in seawater prior to vaccination showed a markedly higher specific mucosal antibody response than sibling fish acclimated in freshwater, regardless of the route of vaccination, whilst the serum antibody response was not affected by salinity. Both mucosal and serum antibodies from fish in seawater and freshwater were capable of binding antigen at salinities similar to full strength seawater in a modified ELISA assay. These results indicate that this euryhaline fish species is riot only able to mount significant specific antibody response in cutaneous mucus, but that these antibodies will function in the marine environment. (c) 2005 Elsevier Ltd. All rights reserved.

Integration of ice-core, marine and terrestrial records for the Australian Last Glacial Maximum and Termination: a contribution from the OZ INTIMATE group

The degree to which Southern Hemisphere climatic changes during the end of the last glacial period and early Holocene (30-8 ka) were influenced or initiated by events occurring in the high latitudes of the Northern Hemisphere is a complex issue. There is conflicting evidence for the degree of hemispheric 'teleconnection' and an unresolved debate as to the principle forcing mechanism(s). The available hypotheses are difficult to test robustly, however, because the few detailed palaeoclimatic records in the Southern Hemisphere are widely dispersed and lack duplication. Here we present climatic and environmental reconstructions from across Australia, a key region of the Southern Hemisphere because of the range of environments it covers and the potentially important role regional atmospheric and oceanic controls play in global climate change. We identify a general scheme of events for the end of the last glacial period and early Holocene but a detailed reconstruction proved problematic. Significant progress in climate quantification and geochronological control is now urgently required to robustly investigate change through this period. Copyright (c) 2006 John Wiley & Sons, Ltd.

Climate change and freshwater zooplankton: what does it boil down to?

Recently, major advances in the climate–zooplankton interface have been made some of which appeared to receive much attention in a broader audience of ecologists as well. In contrast to the marine realm, however, we still lack a more holistic summary of recent knowledge in freshwater. We discuss climate change-related variation in physical and biological attributes of lakes and running waters, high-order ecological functions, and subsequent alteration in zooplankton abundance, phenology, distribution, body size, community structure, life history parameters, and behavior by focusing on community level responses. The adequacy of large-scale climatic indices in ecology has received considerable support and provided a framework for the interpretation of community and species level responses in freshwater zooplankton. Modeling perspectives deserve particular consideration, since this promising stream of ecology is of particular applicability in climate change research owing to the inherently predictive nature of this field. In the future, ecologists should expand their research on species beyond daphnids, should address questions as to how different intrinsic and extrinsic drivers interact, should move beyond correlative approaches toward more mechanistic explanations, and last but not least, should facilitate transfer of biological data both across space and time.