732 resultados para Endosperm weakening


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The Ontong Java Plateau in the western equatorial Pacific contains a deposition record of biserial planktonic foraminifers concentrated in the Paleogene, in which frequencies up to 67% of the planktonic foraminifers are reported, and in the late Neogene, in which a maximum frequency of 48% is reported. Biserial planktonic foraminifers are rare or absent in the latest Oligocene and early Miocene, an interval characterized by warm bottom water and low temperature gradients. These conditions supported a surface assemblage rather than the biserial planktonic foraminifers, whose Neogene species inhabited the oxygen minimum at intermediate depths in the upper water column. Biserial planktonic foraminifers tend to be of high frequency during high sea stands and low frequency during low sea level, presumably in response to the strengthening or weakening of the oxygen minimum. Species extinction and evolution events occur during low sea stands in the Neogene and sometimes correspond to strong reflection horizons of the plateau's seismic stratigraphy. The biserial species are useful biostratigraphic indexes in the plateau section. The last occurrence (LO) of Streptochilus martini corresponds with the Eocene/Oligocene boundary; S. subglobigerum without Neogloboquadrina acostaensis indicates Zone N15; S. latum occurs from the middle of Zone N16 to near the top of Zone N17; S. globigerum ranges from near the top of Zone N17 to the middle of Zone N19/N20; and the S. globulosum continuous range begins just before the first left-to-right coiling change of Pulleniatina, but the species becomes rare in the Pleistocene section.

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Alkenone sea surface temperature (SST) records were generated from the Ocean Drilling Program's (ODP) Sites 1014 and 1016 to examine the response of the California Current System to global climate change during the last 136 ka. The temperature differences between these sites (Delta SST(NEP)=SST(ODP1014)-SST(ODP1016)) reflected the intensity of the California Current and varied between 0.4 and 6.1 °C. A high Delta SST(NEP) (weaker California Current) was found for late marine isotope stage (MIS) 2 and early MIS 5e, while a low Delta SST(NEP) (stronger California Current) was detected for mid-MIS 5e and MIS 1. Spectral analysis indicated that this variation pattern dominated 23- (precession) and 30-ka periods. Comparison of the Delta SST(NEP) and SST based on data from core MD01-2421 at the Japan margin revealed anti-phase variation; the high Delta SST(NEP) (weakening of the California Current) corresponded to the low SST at the Japan margin (the southward displacement of the NW Pacific subarctic boundary), and vice versa. This variation was synchronous with a model prediction of the tropical El Niño-Southern Oscillation behavior. These findings suggest that the intensity of the North Pacific High varied in response to precessional forcing, and also that the response has been linked with the changes of tropical ocean-atmosphere interactions.

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In order to reconstruct hydrographic changes during glacial-interglacial cycles for a transequatorial transect we analyzed oxygen isotopes of Globigerinoides sacculifer (without sac-like chamber) and abundances of Globorotalia truncatulinoides (dextral) from FS Meteor cores GeoB 2204-2 (Brazilian continental slope) and GeoB 1523-1 (Ceara Rise). Delta d18O values of G. sacculifer between the two cores were calculated. Modern Delta d18O (G. sacculifer) is ~0.2 per mill between the two core positions, reflecting differences in sea surface salinity (SSS). Higher SSS at GeoB 1523-1 (Ceara Rise) is the result of increased precipitation in the region of the Intertropical Convergence Zone. During glacials the ?18O records from the two cores converge to the same absolute value, resulting in ??18O values of around 0 per mill. Maximum abundances of G. truncatulinoides (dex) correlate with minimum Delta d18O, suggesting a possible increase of SSS at GeoB 1523-1 during stages 2, 3, 4, and 6, which is related to a glacial weakening of the tropical Hadley Cell [Gates, 1976]. Variations in tropical sea surface temperatures are assumed to be low [Climate: Long-Range Investigation, Mapping, and Prediction (CLIMAP), 1981].

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This paper presents a new fossil pollen record from Tso Moriri (32°54'N, 78°19'E, 4512 m a.s.l.) and seeks to reconstruct changes in mean annual precipitation (MAP) during the last 12,000 years. This high-alpine lake occupies an area of 140 km**2 in a glacial-tectonic valley in the northwestern Himalaya. The region has a cold climate, with a MAP <300 mm, and open vegetation. The hydrology is controlled by the Indian Summer Monsoon (ISM), but winter westerly-associated precipitation also affects the regional water balance. Results indicate that precipitation levels varied significantly during the Holocene. After a rapid increase in MAP, a phase of maximum humidity was reached between ca. 11 to 9.6 cal ka BP, followed by a gradual decline in MAP. This trend parallels the reduction in the Northern Hemisphere summer insolation. Comparison of different palaeoclimate proxy records reveal evidence for a stronger Holocene decrease in precipitation in the northern versus the southern parts of the ISM domain. The long-term trend of ISM weakening is overlaid with several short periods of greater dryness, which are broadly synchronous with the North Atlantic cold spells, suggesting reduced amounts of westerly-associated winter precipitation. Compared to the mid and late Holocene, it appears that westerlies had a greater influence on the western parts of the ISM domain during the early Holocene. During this period, the westerly-associated summer precipitation belt was positioned at Mediterranean latitudes and amplified the ISM-derived precipitation. The Tso Moriri pollen record and moisture reconstructions also suggest that changes in climatic conditions affected the ancient Harappan Civilisation, which flourished in the greater Indus Valley from approximately 5.2 to 3 cal ka BP. The prolonged Holocene trend towards aridity, punctuated by an interval of increased dryness (between ca. 4.5 to 4.3 cal ka BP), may have pushed the Mature Harappan urban settlements (between ca. 4.5 to 3.9 cal ka BP) to develop more efficient agricultural practices to deal with the increasingly acute water shortages. The amplified aridity associated with North Atlantic cooling between ca. 4 to 3.6 and around 3.2 cal ka BP further hindered local agriculture, possibly causing the deurbanisation that occurred from ca. 3.9 cal ka BP and eventual collapse of the Harappan Civilisation between ca. 3.5 to 3 cal ka BP.

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The oceans absorb and store a significant portion of anthropogenic CO2 emissions, but large uncertainties remain in the quantification of this sink. An improved assessment of the present and future oceanic carbon sink is therefore necessary to provide recommendations for long-term global carbon cycle and climate policies. The formation of North Atlantic Deep Water (NADW) is a unique fast track for transporting anthropogenic CO2 into the ocean's interior, making the deep waters rich in anthropogenic carbon. Thus the Atlantic is presently estimated to hold 38% of the oceanic anthropogenic CO2 inventory, although its volume makes up only 25% of the world ocean. Here we analyze the inventory change of anthropogenic CO2 in the Atlantic between 1997 and 2003 and its relationship to NADW formation. For the whole region between 20°S and 65°N the inventory amounts to 32.5 ± 9.5 Petagram carbon (Pg C) in 1997 and increases up to 36.0 ± 10.5 Pg C in 2003. This result is quite similar to earlier studies. Moreover, the overall increase of anthropogenic carbon is in close agreement with the expected change due to rising atmospheric CO2 levels of 1.69% a?1. On the other hand, when considering the subpolar region only, the results demonstrate that the recent weakening in the formation of Labrador Sea Water, a component of NADW, has already led to a decrease of the anthropogenic carbon inventory in this water mass. As a consequence, the overall inventory for the total water column in the western subpolar North Atlantic increased only by 2% between 1997 and 2003, much less than the 11% that would be expected from the increase in atmospheric CO2 levels.

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The Drake Passage (DP) is the major geographic constriction for the Antarctic Circumpolar Current (ACC) and exerts a strong control on the exchange of physical, chemical, and biological properties between the Atlantic, Pacific, and Indian Ocean basins. Resolving changes in the flow of circumpolar water masses through this gateway is, therefore, crucial for advancing our understanding of the Southern Ocean's role in global ocean and climate variability. Here, we reconstruct changes in DP throughflow dynamics over the past 65,000 y based on grain size and geochemical properties of sediment records from the southernmost continental margin of South America. Combined with published sediment records from the Scotia Sea, we argue for a considerable total reduction of DP transport and reveal an up to ~40% decrease in flow speed along the northernmost ACC pathway entering the DP during glacial times. Superimposed on this long-term decrease are high-amplitude, millennial-scale variations, which parallel Southern Ocean and Antarctic temperature patterns. The glacial intervals of strong weakening of the ACC entering the DP imply an enhanced export of northern ACC surface and intermediate waters into the South Pacific Gyre and reduced Pacific-Atlantic exchange through the DP ("cold water route"). We conclude that changes in DP throughflow play a critical role for the global meridional overturning circulation and interbasin exchange in the Southern Ocean, most likely regulated by variations in the westerly wind field and changes in Antarctic sea ice extent.

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Millennial-scale dry events in the Northern Hemisphere monsoon regions during the last Glacial period are commonly attributed to southward shifts of the Intertropical Convergence Zone (ITCZ) associated with an intensification of the northeasterly (NE) trade wind system during intervals of reduced Atlantic meridional overturning circulation (AMOC). Through the use of high-resolution last deglaciation pollen records from the continental slope off Senegal, our data show that one of the longest and most extreme droughts in the western Sahel history, which occurred during the North Atlantic Heinrich Stadial 1 (HS1), displayed a succession of three major phases. These phases progressed from an interval of maximum pollen representation of Saharan elements between ~19 and 17.4 kyr BP indicating the onset of aridity and intensified NE trade winds, followed by a millennial interlude of reduced input of Saharan pollen and increased input of Sahelian pollen, to a final phase between ~16.2 and 15 kyr BP that was characterized by a second maximum of Saharan pollen abundances. This change in the pollen assemblage indicates a mid-HS1 interlude of NE trade wind relaxation, occurring between two distinct trade wind maxima, along with an intensified mid-tropospheric African Easterly Jet (AEJ) indicating a substantial change in West African atmospheric processes. The pollen data thus suggest that although the NE trades have weakened, the Sahel drought remained severe during this time interval. Therefore, a simple strengthening of trade winds and a southward shift of the West African monsoon trough alone cannot fully explain millennial-scale Sahel droughts during periods of AMOC weakening. Instead, we suggest that an intensification of the AEJ is needed to explain the persistence of the drought during HS1. Simulations with the Community Climate System Model indicate that an intensified AEJ during periods of reduced AMOC affected the North African climate by enhancing moisture divergence over the West African realm, thereby extending the Sahel drought for about 4000 years.

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Fine-grained sediment depocenters on continental shelves are of increased scientific interest since they record environmental changes sensitively. A north-south elongated mud depocenter extends along the Senegalese coast in mid-shelf position. Shallow-acoustic profiling was carried out to determine extent, geometry and internal structures of this sedimentary body. In addition, four sediment cores were retrieved with the main aim to identify how paleoclimatic signals and coastal changes have controlled the formation of this mud depocenter. A general paleoclimatic pattern in terms of fluvial input appears to be recorded in this depositional archive. Intervals characterized by high terrigenous input, high sedimentation rates and fine grain sizes occur roughly contemporaneously in all cores and are interpreted as corresponding to intensified river discharge related to more humid conditions in the hinterland. From 2750 to 1900 and from 1000 to 700 cal a BP, wetter conditions are recorded off Senegal, an observation which is in accordance with other records from NW-Africa. Nevertheless, the three employed proxies (sedimentation rate, grain size and elemental distribution) do not always display consistent inter-core patterns. Major differences between the individual core records are attributed to sediment remobilization which was linked to local hydrographic variations as well as reorganizations of the coastal system. The Senegal mud belt is a layered inhomogeneous sedimentary body deposited on an irregular erosive surface. Early Holocene deceleration in the rate of the sea-level rise could have enabled initial mud deposition on the shelf. These favorable conditions for mud deposition occur coevally with a humid period over NW-Africa, thus, high river discharge. Sedimentation started preferentially in the northern areas of the mud belt. During mid-Holocene, a marine incursion led to the formation of an embayment. Afterwards, sedimentation in the north was interrupted in association with a remarkable southward shift in the location of the active depocenter as it is reflected by the sedimentary architecture and confirmed by radiocarbon dates. These sub-recent shifts in depocenters location are caused by migrations of the Senegal River mouth. During late Holocene times, the weakening of river discharge allowed the longshore currents to build up a chain of beach barriers which have forced the river mouth to shift southwards.

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Here we present evidence that the Holocene African monsoon system (AMS) varied in response to the eastern equatorial Atlantic sea-surface temperature (SST). Several short-term episodes of decreased moisture availability as a result of low eastern equatorial Atlantic SST are suggested by planktonic foraminiferal Mg/Ca ratios. These episodes promoted a weakening of the AMS and thus determined the timing and intensity of arid periods. Local sea-surface salinities also reveal regional patterns of precipitation in equatorial western Africa. The high eastern equatorial Atlantic SSTs occur in concert with seasonally increased insolation at low latitudes, suggesting a strong response of African monsoonal precipitation to oceanic conditions at low latitudes.

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We explore the impact of a latitudinal shift in the westerly wind belt over the Southern Ocean on the Atlantic meridional overturning circulation (AMOC) and on the carbon cycle for Last Glacial Maximum background conditions using a state-of-the-art ocean general circulation model. We find that a southward (northward) shift in the westerly winds leads to an intensification (weakening) of no more than 10% of the AMOC. This response of the ocean physics to shifting winds agrees with other studies starting from preindustrial background climate, but the responsible processes are different. In our setup changes in AMOC seemed to be more pulled by upwelling in the south than pushed by downwelling in the north, opposite to what previous studies with different background climate are suggesting. The net effects of the changes in ocean circulation lead to a rise in atmospheric pCO2 of less than 10 atm for both northward and southward shift in the winds. For northward shifted winds the zone of upwelling of carbon- and nutrient-rich waters in the Southern Ocean is expanded, leading to more CO2 outgassing to the atmosphere but also to an enhanced biological pump in the subpolar region. For southward shifted winds the upwelling region contracts around Antarctica, leading to less nutrient export northward and thus a weakening of the biological pump. These model results do not support the idea that shifts in the westerly wind belt play a dominant role in coupling atmospheric CO2 rise and Antarctic temperature during deglaciation suggested by the ice core data.

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The response of the tropical climate in the Indian Ocean realm to abrupt climate change events in the North Atlantic Ocean is contentious. Repositioning of the intertropical convergence zone is thought to have been responsible for changes in tropical hydroclimate during North Atlantic cold spells1, 2, 3, 4, 5, but the dearth of high-resolution records outside the monsoon realm in the Indian Ocean precludes a full understanding of this remote relationship and its underlying mechanisms. Here we show that slowdowns of the Atlantic meridional overturning circulation during Heinrich stadials and the Younger Dryas stadial affected the tropical Indian Ocean hydroclimate through changes to the Hadley circulation including a southward shift in the rising branch (the intertropical convergence zone) and an overall weakening over the southern Indian Ocean. Our results are based on new, high-resolution sea surface temperature and seawater oxygen isotope records of well-dated sedimentary archives from the tropical eastern Indian Ocean for the past 45,000 years, combined with climate model simulations of Atlantic circulation slowdown under Marine Isotope Stages 2 and 3 boundary conditions. Similar conditions in the east and west of the basin rule out a zonal dipole structure as the dominant forcing of the tropical Indian Ocean hydroclimate of millennial-scale events. Results from our simulations and proxy data suggest dry conditions in the northern Indian Ocean realm and wet and warm conditions in the southern realm during North Atlantic cold spells.

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Heinrich events are well documented for the last glaciation, but little is known about their occurrence in older glacial periods of the Pleistocene. Here we report scanning XRF and bulk carbonate d18O results from Integrated Ocean Drilling Program Site U1308 (reoccupation of Deep Sea Drilling Project Site 609) that are used to develop proxy records of ice-rafted detritus (IRD) for the last ~1.4 Ma. Ca/Sr is used as an indicator of IRD layers that are rich in detrital carbonate (i.e., Heinrich layers), whereas Si/Sr reflects layers that are poor in biogenic carbonate and relatively rich in detrital silicate minerals. A pronounced change occurred in the composition and frequency of IRD at ~640 ka during marine isotope stage (MIS) 16, coinciding with the end of the middle Pleistocene transition. At this time, "Hudson Strait" Heinrich layers suddenly appeared in the sedimentary record of Site U1308, and the dominant period of the Si/Sr proxy shifted from 41 ka prior to 640 ka to 100 ka afterward. The onset of Heinrich layers during MIS 16 represents either the initiation of surging of the Laurentide Ice Sheet (LIS) off Hudson Strait or the first time icebergs produced by this process survived the transport to Site U1308. We speculate that ice volume (i.e., thickness) and duration surpassed a critical threshold during MIS 16 and activated the dynamical processes responsible for LIS instability in the region of Hudson Strait. We also observe a strong coupling between IRD proxies and benthic d13C variation at Site U1308 throughout the Pleistocene, supporting a link between iceberg discharge and weakening of thermohaline circulation in the North Atlantic.

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Grain-size records of the terrigenous and calcareous silt fraction, preservation of planktic foraminifera, and benthic foraminiferal stable-isotope data (delta13C, delta18O values of C. wuellerstorfi) at ODP Site 927 on the Ceará Rise (5°27.7'N, 44°28.8'W), are used to reconstruct variations in the history of bottom current strength, ventilation, and carbonate corrosiveness of deep waters during the time interval from 0.8 to 0.3 Ma. Glacial periods are characterized by generally smaller mean sizes of the terrigenous sortable silt fraction (mean(SS)), lower delta13C values, and poorer preservation of planktic foraminifera compared to interglacials. This indicates lower bottom current speeds, larger nutrient contents and more corrosive deep water. By contrast, larger mean(SS) sizes, higher delta13C values, and well preserved planktic foraminifera indicate strong circulation and a well ventilated deep-water mass during interglacials. The observed changes are most likely related to the weakening and strengthening of circulation of Lower North Atlantic Deep Water (LNADW). Cross-spectral analysis between the mean(SS) and benthic delta18O records reveals that minima in mean(SS) occur about 7.6 k.y. after the maximum in ice volume. This indicates a considerable lag time between ice-shield induced changes in LNADW production and subsequent changes in the velocity of LNADW flow in the western equatorial Atlantic. Striking changes in bottom current speed occur regularly during glacial to interglacial transitions. Extremely fine mean(SS) minima point to an almost complete shutdown of bottom current vigor in response to a cessation of LNADW production caused by an enhanced melt water release during the initial phases of deglaciation. However, each of the fine minima extremes is followed by a rapid shift to very high mean(SS) values that indicate strong bottom currents, and hence, vigorous LNADW flow during the early interglacials. After the onset of glacial Stage 12, generally poorer carbonate preservation and higher variability is registered. This coincides with a global decrease in carbonate preservation during the mid-Brunhes (mid-Brunhes dissolution event). Detailed grain-size analysis of the calcareous fine fraction (<63 µm) revealed a considerable reduction of particles in the fraction from 7 to 63 µm during periods of enhanced dissolution. This indicates a preferential dissolution of larger planktic foraminiferal fragments which leads to an enrichment of coccoliths in the calcareous fine fraction.

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Rising levels of atmospheric CO2 lead to acidification of the ocean and alter seawater carbonate chemistry, which can negatively impact calcifying organisms, including mollusks. In estuaries, exposure to elevated CO2 levels often co-occurs with other stressors, such as reduced salinity, which enhances the acidification trend, affects ion and acid-base regulation of estuarine calcifiers and modifies their response to ocean acidification. We studied the interactive effects of salinity and partial pressure of CO2 (PCO2) on biomineralization and energy homeostasis in juveniles of the eastern oyster, Crassostrea virginica, a common estuarine bivalve. Juveniles were exposed for 11 weeks to one of two environmentally relevant salinities (30 or 15 PSU) either at current atmospheric PCO2 (400 µatm, normocapnia) or PCO2 projected by moderate IPCC scenarios for the year 2100 (700-800 µatm, hypercapnia). Exposure of the juvenile oysters to elevated PCO2 and/or low salinity led to a significant increase in mortality, reduction of tissue energy stores (glycogen and lipid) and negative soft tissue growth, indicating energy deficiency. Interestingly, tissue ATP levels were not affected by exposure to changing salinity and PCO2, suggesting that juvenile oysters maintain their cellular energy status at the expense of lipid and glycogen stores. At the same time, no compensatory upregulation of carbonic anhydrase activity was found under the conditions of low salinity and high PCO2. Metabolic profiling using magnetic resonance spectroscopy revealed altered metabolite status following low salinity exposure; specifically, acetate levels were lower in hypercapnic than in normocapnic individuals at low salinity. Combined exposure to hypercapnia and low salinity negatively affected mechanical properties of shells of the juveniles, resulting in reduced hardness and fracture resistance. Thus, our data suggest that the combined effects of elevated PCO2 and fluctuating salinity may jeopardize the survival of eastern oysters because of weakening of their shells and increased energy consumption.