Biostratigraphy and paleobiogeographic affinities of the Jurassic brachiopod assemblages from Sierra Espuña (Maláguide Complex, Internal Betic Zones, Spain)

The assemblages of Early Jurassic brachiopods (Pliensbachian - Toarcian) from Sierra Espuña (Murcia Province, SE Spain) are described. This is the only area in the Internal Zones of the Betic Cordillera, corresponding to the margins of the Alborán Terrane, where Jurassic brachiopods are known to occur. In the tectonic Unit of Morrón de Totana (more southward located) assemblage MT1 of Late Pliensbachian age has been characterized. This assemblage has been subdivided into three successive sub-assemblages: MT1a (Algovianum Zone), MT1b (Emaciatum Zone, Solare Subzone) and MT1c (Emaciatum Zone, Elisa Subzone). Northward, in the Perona tectonic Unit two distinct assemblages, P1 (Latest Sinemurian - Early Pliensbachian) and P2 (Early Toarcian, Serpentinum Zone) have been recognized. Differences between the assemblages from the two tectonic units are evident after the paleobiogeographical analysis. In the Morrón de Totana Unit, taxa with Mediterranean affinities occur. MT1 assemblage is very similar to assemblages previously known in the Eastern Subbetic as well as in other areas of the Mediterranean Province. In the Perona Unit the Mediterranean affinity of the assemblages is not so evident. P1 Assemblage consists of widely distributed taxa, lacking in the most characteristic elements of the Mediterranean Province which, however, are present in neighbouring Betic areas. P2 Assemblage belongs to the Spanish Province that develops in Western Tethys after the Early Toarcian Mass Extinction Event. The occurrence in this assemblage of Prionorhynchia aff. msougari Rousselle, until now only found in North Africa, indicates a closer connection of the Perona Unit with the African paleomargin of the Tethys than with the South Iberian paleomargin. The paleobiogeographical data suggest a more southern and marginal (close to epicontinental areas) position of the Perona Unit than the Morrón de Totana Unit.

Evolution of the last koninckinids (Athyridida, Koninckinidae), a precursor signal of the early Toarcian mass extinction event in the Western Tethys

Koninckinids are a suitable group to shed light on the biotic crisis suffered by brachiopod fauna in the Early Jurassic. Koninckinid fauna recorded in the late Pliensbachian–early Toarcian from the easternmost Subbetic basin is analyzed and identified as a precursor signal for one of the most conspicuous mass extinction events of the Phylum Brachiopoda, a multi-phased interval with episodes of changing environmental conditions, whose onset can be detected from the Elisa–Mirabile subzones up to the early Toarcian extinction boundary in the lowermost Serpentinum Zone (T-OAE). The koninckinid fauna had a previously well-established migration pattern from the intra-Tethyan to the NW-European basins but a first phase with a progressive warming episode in the Pliensbachian–Toarcian transition triggered a koninckinid fauna exodus from the eastern/central Tethys toward the westernmost Mediterranean margins. A second stage shows an adaptive response to more adverse conditions in the westernmost Tethyan margins and finally, an escape and extinction phase is detected in the Atlantic areas from the mid-Polymorphum Zone onwards up to their global extinction in the lowermost Serpentinum Zone. This migration pattern is independent of the paleogeographic bioprovinciality and is unrelated to a facies-controlled pattern. The anoxic/suboxic environmental conditions should only be considered as a minor factor of partial control since well-oxygenated habitats are noted in the intra-Tethyan basins and this factor is noticeable only in the second westward migratory stage (with dwarf taxa and oligotypical assemblages). The analysis of cold-seep proxies in the Subbetic deposits suggests a radiation that is independent of methane releases in the Subbetic basin.

Braquiópodos del Jurásico Inferior del norte de España en la colección Jiménez de Cisneros

Se actualizan sistemáticamente las especies de braquiópodos del Jurásico Inferior procedentes de varias localidades del N y NE de España alojados en la colección Jiménez de Cisneros, refiriéndolas a contextos actualmente contrastados, dotándolas así de un conveniente marco lito- y cronoestratigráfico. Se encuentran representados especímenes de la Rama Castellana y Aragonesa de la Cordillera Ibérica, Asturias, Pirineos y la Cordillera costera Catalana. La actualización ha permitido identificar 18 especies distintas desde el Sinemuriense al Aaleniense basal, contextualizándolas y comparándolas paleobiogeográficamente con el conjunto de plataformas que ocupaban los paleomárgenes de la Subplaca Ibérica durante el Jurásico Inferior.

Brachiopod assemblages from the Early-Middle Jurassic transition in the Eastern Subbetic (SE Spain): Systematic and palaeobiogeographic implications and palaeoenvironmental significance

New data on brachiopod assemblages recorded in the Eastern Subbetic area (Alicante, SE Spain) and attributed to the Early Bajocian (Humphriesianum Zone and/or immediately older) are provided. Eleven species have been distinguished and reported for the first time in the Subbetic domain of the Betic Cordillera. The description of the morphological evidences on each analysed taxa, especially in relation to their internal morphology, brings new implications in the systematics of the Middle Jurassic brachiopods. The analysis of faunistic affinity between the recorded assemblages and those from other palaeogeographic domains, shows that the Subbetic brachiopod fauna has a clear Mediterranean affinity, as proved by the different species belonging to the genera Striirhynchia, Septocrurella, Mondegia?, Karadagithyris, Linguithyris, Papodina?, Viallithyris?, and Zugmayeria?. It is also evidenced that the Early-Middle Jurassic transition in the Eastern Subbetic accounted, in qualitative terms, a remarkable interval of faunistic renewal in the brachiopod assemblages, strongly influenced by a complex tectonic and stratigraphic framework controlled by a period of intense extensional tectonics, globally framed in the evolution of the Atlantic Ocean.

New theropod remains from the Tiourarén Formation (?Middle Jurassic, Niger) and their bearing on the dental evolution in basal tetanurans

A fragment of a maxilla and isolated theropod teeth from the (?) Middle Jurassic Tiourarén Formation are described. The specimens come from Tadibene, in the rural community of Aderbissinat, Thirozerine Department, Agadez Region, Niger. They were identified through direct comparison with teeth previously described in the literature as well as on the basis of discriminant and morphometric analyses. Our results suggest they belong to Ceratosauridae, Megalosauridae, and the oldest representatives of Spinosauridae. The analyzed sample shows some uncommon characters, such as spinosaurid-like ornamentation in megalosaurid-like teeth, or spinosaurid-like teeth with a low number of denticles, which sheds light on tooth morphology and dental evolution in basal tetanurans and early spinosaurids.