36 resultados para upland steppes
Resumo:
This paper presents results of 2 years (from January 2005 to December 2006) of measurement of N2O fluxes from the native and grazed Leymus chinensis (LC) steppes in Inner Mongolia, China using the static opaque chamber method. The measurement was at a frequency of twice per month in the growing season and once per month in the non-growing season. In addition, the possible effect of water-heat factors on N2O fluxes was statistically analyzed. The results indicated that there were distinct seasonal patterns in N2O fluxes with large fluxes in spring, summer, and autumn but negative fluxes in winter. The annual net emission of N2O ranging from 0.24 to 0.30 kg N2O-N ha(-1) and from 0.06 to 0.26 kg N2O-N ha(-1) from the native and grazed LC steppe, respectively. Grazing activities suppressed N2O production. In the growing season, soil moisture was the primary driving factor of N2O fluxes. The high seasonal variation of N2O fluxes was regulated by the distribution of effective rainfall, rather than precipitation intensity. Air temperature or soil temperature at 0, 5, and 10 cm depth was the most restricting factor of N2O fluxes in the non-growing season.
Resumo:
We measured the stable carbon isotope ratios for muscle of the upland buzzards (Buteo hemilasius), plateau pika (Ochotoma curzoniae), root vole (Microtus oeconomus), plateau zokor (Myospalax fontanierii) and passerine bird species at the Haibei Alpine Meadow Ecosystem Research Station (HAMERS), and provided diet information of upland buzzards with the measurement of stable carbon isotopes in tissues of these consumers. The results showed that δ~(13)C values of small mammals and passerine bird species ranged from -25.57‰ to -25.78‰ (n = 12), and from -24.81‰ to -22.51% (n = 43), respectively, δ~(13)C values of the upland buzzards ranged from -22.60‰ to -23.10‰ when food was not available. The difference in δ~(13)C values (2.88‰±0.31‰) between upland buzzards and small mammals was much larger than the differences reported previously, 1‰-2‰, and showed significant difference, while 1.31‰±0.34‰ between upland buzzard and passerine bird species did not differ from the previously reported trophic fractionation difference of 1‰-2‰. Estimation of trophic position indicated that upland buzzards stand at trophic position 4.23, far from that of small mammals, i.e., upland buzzards scarcely captured small mammals as food at the duration of food shortage. According to isotope mass balance model, small mammals contributed 7.89% to 35.04% of carbon to the food source of the upland buzzards, while passerine bird species contributed 64.96% to 92.11%. Upland buzzards turned to passerine bird species as food during times of shortage of small mammals. δ~(13)C value, a useful indicator of diet, indicates that the upland buzzards feed mainly on passerine bird species rather than small mammals due to "you are what you eat" when small mammal preys are becoming scarce.
Resumo:
Summer diets of two sympatric raptors Upland Buzzards (Buteo hemilasius Temminck et Schlegel) and Eurasian Eagle Owls (Bubo bubo L. subsp. Hemachalana Hume) were studied in an alpine meadow (3250 m a.s.l.) on Qinghai-Tibet Plateau, China. Root voles Microtus oeconomus Pallas, plateau pikas Ochotona curzoniae Hodgson, Gansu pikas O. cansus Lyon and plateau zokors Myospalax baileyi Thomas were the main diet components of Upland Buzzards as identified through the pellets analysis with the frequency of 57, 20, 19 and 4%, respectively. The four rodent species also were the main diet components of Eurasian Eagle Owls basing on the pellets and prey leftovers analysis with the frequency of 53, 26, 13 and 5%, respectively. The food niche breadth indexes of Upland Buzzards and Eurasian Eagle Owls were 1.60 and 1.77 respectively (higher value of the index means the food niche of the raptor is broader), and the diet overlap index of the two raptors was larger (C-ue = 0.90) (the index range from 0 - no overlap - to I - complete overlap). It means that the diets of Upland Buzzards and Eurasian Eagle Owls were similar (Two Related Samples Test, Z = -0.752, P = 0.452). The classical resource partitioning theory can not explain the coexistence of Upland Buzzards and Eurasian Eagle Owls in alpine meadows of Qinghai-Tibet Plateau. However, differences in body size, predation mode and activity rhythm between Upland Buzzards and Eurasian Eagle Owls may explain the coexistence of these two sympatric raptors.
Resumo:
To clarify the response of soil organic carbon (SOC) content to season-long grazing in the semiarid typical steppes of Inner Mongolia, we examined the aboveground biomass and SOC in both grazing (G-site) and no grazing (NG-site) sites in two typical steppes dominated by Leymus chinensis and Stipa grandis, as well as one seriously degraded L. chinensis grassland dominated by Artemisia frigida. The NG-sites had been fenced for 20 years in L. chinensis and S. grandis grasslands and for 10 years in A. frigida grassland. Above-ground biomass at G-sites was 21-35% of that at NG-sites in L. chinensis and S. grandis grasslands. The SOC, however, showed no significant difference between G-site and NG-site in both grasslands. In the NG-sites, aboveground biomass was significantly lower in A. frigida grassland than in the other two grasslands. The SOC in A. frigida grassland was about 70% of that in L. chinensis grassland. In A. frigida grassland, aboveground biomass in the G-site was 68-82% of that in the NG-site, whereas SOC was significantly lower in the G-site than in the NG-site. Grazing elevated the surface soil pH in L. chinensis and A. frigida communities. A spatial heterogeneity in SOC and pH in the topsoil was not detected the G-site within the minimal sampling distance of 10 m. The results suggested that compensatory growth may account for the relative stability of SOC in G-sites in typical steppes. The SOC was sensitive to heavy grazing and difficult to recover after a significant decline caused by overgrazing in semiarid steppes.
Resumo:
We measured the stable carbon and nitrogen isotope ratios for muscles of the upland buzzards (Buteo hemilasius) and their potential food sources, plateau pikas (Ochotona curzoniae), Qinghai voles (Lasiopodomys fuscus), plateau zokors (Myospalax fontanierii), and several passerine bird species at the alpine meadow in Maduo county, Guoluo prefecture of Qinghai province, People's Republic of China, to provide diet information of upland buzzards, highlighting different diet composition of upland buzzards exposed to different locations. The results demonstrated that stable carbon isotope ratios of upland buzzards, passerine birds, plateau pikas, plateau zokors, and Qinghai voles were -24.42 +/- 0.25parts per thousand, -22.89 +/- 1.48parts per thousand, -25.30 +/- 1.47parts per thousand, -25.78 +/- 0.22parts per thousand, and -25.41 +/- 0.01parts per thousand, respectively, and stable nitrogen isotope ratios were 7.89 +/- 0.38parts per thousand, 8.37 +/- 2.05parts per thousand, 5.83 +/- 1.10parts per thousand, 5.23 +/- 0.34parts per thousand, and 8.86 +/- 0.06parts per thousand, respectively. Fractionation of stable carbon and nitrogen isotope ratios between upland buzzards and their food were 1.03parts per thousand and 2.11parts per thousand, respectively. Based on mass balance principle of stable isotopes and the Euclidean distance mixing model, upland buzzards depended mainly on plateau pikas as food (74.56%). Plateau zokors, Qinghai voles, and passerine birds only contributed a small proportion (25.44%) to diets of upland buzzards. The results were closely accordant with analyses of stomach contents and food pellets, which firmly supported the feasibility of using stable carbon and nitrogen isotope ratios to investigate diet information of upland buzzards. Another study based on stable carbon isotopes showed that upland buzzards living in the Haibei prefecture (another prefecture located in the southeast Qinghai province) mainly preyed on passerine birds (64.96% or more) as food supply. We were alarmed by the preliminary results that widespread poisoning activities of small mammals could reshape the food composition of upland buzzards, influencing the stability and sustainability of the alpine meadow. Bio-control on rodent pests should be carried out rather than the chemical measures.
Resumo:
1.以MS无机盐+B 5有机成分为基本培养基,附加2,4-D,KT,ZT等激素,以农艺性状良好,抗枯萎病的棉花品种“冀492”为材料,建立了愈伤组织诱导,体细胞胚胎发生和植株再生体系. 2.实验结果表明,2,4-D是诱导愈伤组织产生的促进因子,效果明显好于其它生长素类物质,但是随着培养基中2,4-D浓度的升高,愈伤组织状态也会变差,褐化严重,而且,附加O.lppm 2,4-D,O.lppm K T的培养基是较为合适的培养基.IAA,NAA对愈伤组织的诱导不十分理想,此外,愈伤组织的产生还同外植体,基因型和一些物理条件有关, 胚性愈伤组织的产生,需不断调节培养基中激素浓度,逐步降低2,4 -D浓度,并添加低浓度ZT,胚状体要在去除2,4-D的培养基上才能产生.虽然,2,4-D对于胚性愈伤组织的诱导是必须的,但是却抑制胚状体的发育. 3.培养基中硝态氮和氨态氮比例,激素浓度和种类,活性炭的添加,对于胚状体的萌发和成熟有重要影响,硝酸钾加倍,硝酸铵减半,并附加低浓度ABA和活性炭是胚状体诱导和成苗(去除ABA)的适宜培养基,而附加0.2ppm N A A,0.2ppm ZT和活性炭的M S2培养基是胚状体成熟的适宜培养基.通过对胚性愈伤组织的干燥处理,培养基中低浓度ABA和活性炭的添加,比较有效地抑制了畸形胚的产生,大大地提高了正常胚的比例. 4.对陆地棉“冀4 9 2”的下胚轴和子叶利用含Bt抗虫基因的根农杆菌(AgrobacterIIm tumefrens)进行了遗传转化研究,在筛选培养基上获得了大量生长旺盛的阳性愈伤组织,葡糖酸苷检测结果表明,大部分愈伤组织均有G us基因的表达. 5.通过花粉管通道法,进行了将含有Bt抗虫基因DNA的大肠杆菌质粒导入陆地棉“冀492”的实验,并收获了近1 0,000粒种子,为从大量种子中筛选出抗性种质材料,建立了卡那霉素田间初步筛选法. 6.利用活体压片和石蜡切片技术对体细胞胚胎发生过程和胚状体起源进行了观察,发现胚状体发生主要是通过类合子途径进行的,胚状体可能起源于单细胞. 7.通过对相同培养基上的非胚性愈伤组织,胚性愈伤组织和不同时期胚状体的可溶性蛋白SDS - PAGE电泳分析,发现16KD和50K D蛋白特异性地存在于胚性愈伤和各期胚状体中,该蛋白可作为胚性愈伤组织的筛选标记.对不同发育时期的愈伤组织进行了同工酶和可溶性蛋白的IE F/SD S-PA G E双向电泳分析,结果表明,不同发育时期的愈伤组织同工酶酶谱和可溶性蛋白电泳图谱之间具有较大的差别,分化前期的胚性愈伤组织酶的活性强,种类多,并且有新的蛋白斑点的出现,这些都可能与体细胞胚胎发生相关.
Resumo:
本文通过根农杆菌(Agrobacterium tumfaciens)介导法分别将Signal和KDEL修饰的豇豆胰蛋白酶抑制剂(Cowpea trypsin inhibitor, CpTI)基因、豌豆外源凝集素(Pea lectin, P-Lec)和大豆Kunitz型胰蛋白酶抑制剂(Soybean Kunitz typsin inhibitor, SKTI)双价抗虫基因、雪花莲外源凝集素(Galanthus nivals agglutinin, GNA)基因以及高效复合启动子OM控制的苏云金杆菌(Bacillus thuringiensis, B.t.)杀虫毒蛋白基因导入了陆地棉(Gossypium hirsutum L.)栽培品种新陆早1号、新陆中2号、晋棉7号、冀合321、辽9和晋棉12号,并获得了大批转基因再生植株。 实验中对影响棉花转化和再生的一些条件进行了研究,从根农杆菌培养、棉花无菌苗的制备、转化操作和共培养等方面对转化条件进行了探讨;从激素配化、植物表达载体、外植体类型、基因型等方面对抗性愈伤组织的诱导进行了摸索;从激素、从碳源、培养容器、pH值、抗褐化剂及固化剂的选择等方面对影响植株再生的条件进行了优化。 本文开创性地采用嫁接代替移栽,从而极大地提高了转化植株定植成活率,缩短了缓苗时间并增加了转化植株当代的繁殖系数。 在建立了一套较为高效的陆地棉转化及再生系统基础上,本文还进行了其它转化方式和转化体系的初步探讨。利用棉花幼嫩种子无菌苗下胚轴作为外植体,通过改变愈伤组织诱导培养基配方面提高胚性愈伤组织的诱导频率,进而得到更多的体细胞胚状和再生植株,缩短再生周期;尝试用胚性愈伤组织作为外植体的根农杆菌介导法转化,确定了一些与转化有关的条件;建立了一套棉花茎尖培养程序,为运用基因枪法轰击棉花茎尖分生组织或用根农杆菌直接转化茎尖分生组织,以克服根农杆菌转化棉花时体胚发生的基因型局限开辟了一条新途径。 本文还建立了一种快速鉴定转化植株后代的方法。这一简便方法还有助于进行转基因棉纯合系的筛选以及外源基因的遗传稳定性研究。 转基因植株经Npt-II ELISA、PCR、PCR Southern 检测证明外源抗虫基因CpTI、SKTI、P-lec、GNA以及B.t.基因已存在于转化植株基因组内。修饰的CpTI转基因植株抗棉铃虫(Heliothis armigera Hubner)试验结果表明,其杀虫效果显著优于前期未修饰的CpTI转化植株。P-lec和SKTI双价转基因植株抗棉铃虫试验结果表明,转基因植株对棉铃虫幼虫具有较强的杀虫活力。 目前,已获得转以上抗虫基因棉花T1代植株。为今后进一步将植物基因工程技术应用于棉花遗传改良打下了基础。
Resumo:
本文采用野外观测和室内测定相结合的方法,研究了内蒙古草原两种主要的土地利用方式(开垦和放牧)对土壤碳库和温室气体通量的影响,结果如下: 1.内蒙古草甸草原开垦后30年后,与天然草原相比,在休闲年份0~20 cm土壤有机碳和土壤微生物量碳含量均没有显著下降,但开垦导致土壤易分解碳下降了24%。土壤易氧化碳受季节性水热因子的驱动,表现出明显的季节变化动态。因此,土壤易分解碳的较土壤碳库的其他组分对开垦更敏感,是表征土壤管理措施引起有机质变化的一个重要指标。 草原开垦后,土壤-植物系统氧化大气甲烷的能力明显提高,农田和天然草原CH4平均吸收通量分别是48.9 和 29.0 μg C m-2 h-1,开垦后增加了1.7倍。开垦没有改变CH4吸收通量 “夏季高秋季低” 的季节变化规律(由降水决定);开垦使N2O的平均释放通量增加了47%,农田和天然草原N2O平均吸收通量分别是56.6 和 38.6 ugN m-2 h-1;开垦同时也增大了通量的变异幅度;但没有改变N2O季节变化规律,只是出现高峰的时间较天然草原推后约10天左右。 2.开垦后的农田土壤在模拟添加厩肥后,刺激了土壤微生物的呼吸代谢,使CO2的释放量增加了5-7倍。试验期间总体排放的CO2中,约60%来源于羊粪,40% 来源于土壤。两种土壤即羊草顶级草原土壤(高碳高氮)和冷蒿-小禾草退化草原土壤(低碳低氮),在CO2的释放总量和释放比例上都没有显著性差异。添加厩肥均造成两种土壤碳库的净碳损失,并且退化草原土壤(7.0%)的土壤净碳损失要大于羊草草原(2.6%)。说明与开垦后的高C土壤相比,在已经退化草原的低C土壤上施厩肥将趋向于土壤更大的净碳损失。 3.自由放牧22年后,羊草草原0~10 cm土壤有机碳、微生物量碳和易分解碳分别下降了14.1%、27.9%和22.0%;大针茅草原0~5 cm土壤有机碳和微生物量碳分别下降了27.6%和38.2%。两类草原土壤碳组分的季节变化受水热因子的驱动,大针茅草原季节波动出现高峰的时间较羊草草原迟。土壤微生物量碳在表征羊草草原和大针茅草原土壤碳素的动态变化时,要敏感于土壤总有机碳。放牧对冷蒿-小禾草草原土壤各碳素组分影响不明显。在表征放牧对冷蒿-小禾草草原土壤的影响指示上,MB-C/ Org-C和Lab-C/ Org-C要比MB-C和Lab-C更加敏感。这说明在研究放牧对草原土壤碳库影响时,不同的草原类型应使用不同的指标来表征其变化。 内蒙古羊草草原是大气CH4的汇,自由放牧增加土壤对CH4的吸收。CH4平均吸收通量增加了27%,但CH4吸收的季节变化形式没有改变;放牧使。自由放牧还增大了N2O的排放通量,将原来N2O源、汇的双重功能改变为单一的源功能;放牧使N2O平均释放通量增加了1倍;放牧显著增加了羊草草原向大气排放CO2的量(p<0.05),并且年度排放量范围也有所增大。 4.草原羊尿斑土壤的pH和NH4+浓度在施后显著升高,但土壤微生物C和N没有明显变化,尿斑N素会发生大量的流失。粪斑和厩肥斑中各有46%和27%的N素分解后转移到植物中。羊草种群斑块上粪尿斑引起CO2和N2O通量的变化,要大于星毛委陵菜种群斑块。与植被类型的影响相比,羊粪尿斑尤其是尿斑对温室气体通量的影响更大。尿斑既降低了土壤对CH4的吸收,又增加了CO2和N2O的释放,使粪尿斑上相当于CO2的净排放量比对照土壤增加了15%。 在内蒙古草原中等放牧条件下,家畜粪尿斑在放牧草地上的覆盖面积约只有2%,与未被家畜排泄物覆盖的草原土壤 相比,粪尿斑对内蒙古草原温室气体总体收支产生的影响可以忽略不计。因此内蒙古草原地区温室气体减排措施的重点,应放在家畜的食性食量对温室气体的影响以及厩肥的科学利用上。但随放牧强度的加大,家畜排泄物覆盖草地的面积将大大增加,加之放牧生态系统中家畜瘤胃代谢产生的的大量温室气体,其对草原温室气体的核算将会产生的影响也是不容忽视的。
Resumo:
地上净初级生产力(ANPP)是陆地生态系统碳循环的重要组成部分,但由于估测ANPP的方法不同使得对ANPP的估测值存在很大的不确定性。本文采用3种方法(群落中所有种群当年最大地上生物量之和(ANPPC1)、当年群落最大地上生物量(ANPPC2)以及每年固定日期(8月30日)的地上生物量(ANPPC3))在种群、功能群和群落水平上分别对连续19年(1980~1998)的内蒙古羊草和大针茅草原生态系统功能(如ANPP、植物多样性和水分利用效率(WUE))的动态变化进行了比较分析,同时探讨了不同估测方法下气候和放牧对ANPP的影响,在此基础上利用DNDC模型进行了ANPP的模拟和敏感性分析研究,主要结果包括: 在羊草和大针茅草原群落中,不同主要植物种群或功能群多年平均地上最大生物量出现的时间不同,而同一植物种群或功能群的年地上最大生物量出现的时间存在年际间的变化。采用当年最大地上生物量和8月30日固定日期的地上生物量作为群落ANPP的这两种方法高估了建群种或禾草功能群在群落中的作用。 羊草草原群落多年平均ANPPC1、ANPPC2和 ANPPC3分别是257.5、190.1和166.0 g.m-2;相应的大针茅草原多年平均ANPPC1、ANPPC2和 ANPPC3分别是180.4、132.8和122.5 g.m-2。就群落生产力而言,后两种常用的方法二和方法三分别低估了草原群落ANPP 14.2%~40.0%和15.5~59.0%。本文研究表明尽管ANPPC1与ANPPC2和ANPPC3之间存在显著的差异,但二者之间存在极显著的相关性:羊草草原的ANPPC1= 59.587+1.061×ANPPC2(r2=0.865, p<0.001),ANPPC1= 92.329+1.017×ANPPC3(r2=0.569, p<0.001);大针茅草原的ANPPC1= 32.918+1.114×ANPPC2(r2=0.814, p<0.001),ANPPC1= 76.120+0.875×ANPPC3(r2=0.499, p=0.001)。 种群、功能群和群落地上净初级生产力与气候因子间的关系因不同的估测方法而异。羊草草原的建群种羊草种群仅ANPPS3与8月和11月份的平均温度间存在显著相关性,而大针茅草原群落的建群种大针茅种群的ANPPS1和ANPPS2与3月份平均最高气温间呈负相关关系。在羊草草原群落,杂类草功能群ANPPF1和ANPPF2与3月份气温呈负相关关系;而灌木半灌木功能群ANPPF1和ANPPF2与3月份降水呈负相关关系。在大针茅草原群落,禾草功能群ANPPF1与5月份最高平均气温呈显著负相关关系。羊草草原群落ANPPC1和ANPPC3分别与11月份最低温度和平均温度存在显著负相关关系;大针茅草原群落ANPPC1与2和6月份月降水量间相关性显著,ANPP2与5月和11月份月平均最低温度呈显著负相关关系,而4~9月份,1和4月份平均最低温度对群落ANPPC3起决定作用。 在羊草和大针茅草原群落,由方法一得到的群落水分利用效率、Shannon植物多样性指数和均匀度指数与方法二或方法三得到的相应的指数间存在显著的差异,方法二和方法三得到的值间差异不显著。群落地上净初级生产力与相应估测方法的植物Shannon多样性指数和均匀度指数之间相关性不显著。 放牧条件下羊草或大针茅草原群落中的建群种羊草和大针茅在群落中的相对地上生物量较围栏内的相应植物种群的降低,而糙隐子草种群在群落中的比例上升。不同的放牧管理条件下,群落中植物种群地上现存量的季节动态发生变化。群落的植物组成及植物种群地上生物量在群落中总生物量的比例发生了明显的变化。利用遥感来估测地上净初级生产力时,分别低估了羊草和大针茅草原ANPP 52%和27%。 DNDC模型可以很好地模拟内蒙古典型草原生态系统的地上生物量,通过敏感性分析表明降水是草原植物生长的主要限制因子,在降水量增加或降低至日降水的30%时,模拟的地上生物量显著地高于或低于实测地上生物量值。在多年平均降水量为347mm的情况下,随着土壤粘粒含量的增加,地上生物量逐渐降低,与北美草原一致。
Resumo:
Manu National Park of southern Peru is one of the most renowned protected areas in the world, yet large-bodied vertebrate surveys conducted to date have been restricted to Cocha Cashu Biological Station, a research station covering <0.06 percent of the 1.7Mha park. Manu Park is occupied by >460 settled Matsigenka Amerindians, 300-400 isolated Matsigenka, and several, little-known groups of isolated hunter-gatherers, yet the impact of these native Amazonians on game vertebrate populations within the park remains poorly understood. On the basis of 1495 km of standardized line-transect censuses, we present density and biomass estimates for 23 mammal, bird, and reptile species for seven lowland and upland forest sites in Manu Park, including Cocha Cashu. We compare these estimates between hunted and nonhunted sites within Manu Park, and with other Neotropical forest sites. Manu Park safeguards some of the most species-rich and highest biomass assemblages of arboreal and terrestrial mammals ever recorded in Neotropical forests, most likely because of its direct Andean influence and high levels of soil fertility. Relative to Barro Colorado Island, seed predators and arboreal folivores in Manu are rare, and generalist frugivores specializing on mature fruit pulp are abundant. The impact of such a qualitative shift in the vertebrate community on the dynamics of plant regeneration, and therefore, on our understanding of tropical plant ecology, must be profound. Despite a number of external threats, Manu Park continues to serve as a baseline against which other Neotropical forests can be gauged.
Resumo:
Nostoc flagelliforme, which is distributed in arid or semiarid steppes of the west and west-northern parts of China, has been used by the Chinese as a food delicacy and for its herbal values for hundreds of years. However, the resource is being over-exploited and is diminishing, while the market demands are increasing with the economic growth. This review deals mainly with the Chinese studies on the ecology, physiology, reproduction, morphology and culture of this species in an attempt to promote research and development of its cultivation technology.
Resumo:
通过对长武王东沟村农户土地利用结构及经济收入的详细调查,揭示了2007年王东沟小流域不同收入类型农户的土地利用结构现状、1998-2007年的土地利用结构变化情况、产业结构和投入产出情况及其经济效益等。结果表明当前农户最主要的土地利用方式是果园和耕地,收入的主要来源是工副业和果业,各类型间农户收入及其构成差异显著。不同收入水平农户的效益差异与有自然条件、管理水平及劳动力水平等因素有关。提出王东沟小流域应结合目前政策形势调整农户种植结构,合理配置土地资源,发展第三产业以提高经济收入等建议。
Resumo:
沙蜥属Phrynocephalus Kaup,1825隶属于爬行纲(Reptilia)有鳞目(Squamata)蜥蜴亚目(Lacertilia)鬣蜥科(Agamidae),是欧亚大陆荒漠和稀疏草原常见蜥蜴。沙蜥属的分类及系统演化关系、地理分布格局与新生代第三纪以来古地中海的变迁、青藏高原的抬升及亚洲内陆干旱荒漠化的过程有密切的关系,长期以来有关沙蜥属的研究一直受到中外学者们的关注。由于沙蜥属地理分布广、形态变异大、体色和斑纹变化复杂,虽然前人使用过许多形态性状来描述和分类沙蜥属物种,但是仍然存在许多问题。性状的分类学意义不明确是造成这些问题的主要原因之一,因此本研究针对沙蜥属常用的鉴别性状进行分类意义的分析,希望能对沙蜥属物种鉴定及分类学其它研究有所裨益。 中国沙蜥属物种主要分布于西北的干旱荒漠区域及青藏高原的大部分地区,大约为18种。 本文研究了中国境内12种沙蜥:青海沙蜥(Phrynocephalus vlangalii)、西藏沙蜥(P. theobaldi)、南疆沙蜥(P. forsythii)、变色沙蜥(P. versicolor)、旱地沙蜥(P. helioscopus)、荒漠沙蜥(P. przewalskii)、乌拉尔沙蜥(P. guttatus)、草原沙蜥(P. frontalis)、叶城沙蜥(P. axillaris)、白稍沙蜥(P. koslowi)、无斑沙蜥(P. immaculatus)和白条沙蜥(P. albolineatus),对它们的65项外部形态性状进行了观察和测量,其中数量性状29项、质量性状36项。评价了这些性状的序级性、间断性和代表性,结论如下: 1. 对于数量性状,得出了适合各级分类的数值区间; 2. 给出了在不同序级上适合分类的质量性状。 并利用各性状评价的结果,给出12种沙蜥的检索表,以及对中国沙蜥物种某些尚存在争议的问题进行了探讨。 详细记录了青海沙蜥红原亚种的骨骼系统,首次发现并命名了肘骨(elbow bone)和垫骨(stepping bone),为沙蜥属系统学研究补充了骨骼方面的证据;解剖了乌拉尔沙蜥、旱地沙蜥、荒漠沙蜥的雌体和雄体的骨骼系统,并在14项骨骼形态性状上对这3种沙蜥进行了比较。 Phrynocephalus (Squamata,Agamidae) is a familiar genus of lizards inhabited desert and sparse steppes in Eurasia. The taxonomics, phylogenetics and distribution pattern of Phrynocephalus are relative intensely to these events: the vicissitudes of the archaic Mediterranean sea since the Cainozoic, the uplift of Qingzang Plateau and the expending arid areas in the inland of Asia. Owing to the wide distribution, the large variability of the morphology and the different colors in Phrynocephalus, it is difficult to identify them. Tough many morphological characters are used to describe and discriminate them,a lot of questions still exist. One of the most important reasons is the confusion in the morphological characters. In this study, we demonstrate the validity and the invalidity of the familiar characters. There are about 18 species of the genus Phrynocephalus in China, which exist in arid desert in Northwest China and Qingzang Plateau. Twelve Chinese species was analyzed in this paper. They are P. vlangalii,P. theobaldi,P. forsythia,P. versicolor,P. helioscopus,P. przewalskii,P. guttatus,P. frontalis,P. axillaris,P. koslowi,P. immaculatus,P. albolineatus. We measure 29 quantitative characters and observe 36 qualitative characters in each individual. Through analyzing these characters, we made some conclusions as follows: 1. to every quantitative character, we get a clear numeric area to discriminate the different operational taxonomic units. 2. we chose the valid qualitative characters in these operational taxonomic units. This paper is the first to describe the “elbow bone”, which is a bone in pectoral appendage equivalent to patella,and “stepping bone”, which is a bone under carpal. A detailed description of the skeletal system of female Phrynocephalus vlangalii hongyuanensis was conducted. We also anatomise the skeletal systems of three species: P. guttatus,P. przewalskii,P. helioscopus, and compare or contrast 14 skeletal characters in them. What’s more,this paper offers some suggestions to the questions of Chinese Phrynocephalus species and keys to 12 species of Phrynocephalus basing on our conclusions on the evaluation of the morphological characters.
Resumo:
以陆地棉 (Gossypium hirsutum L.Zhongmain No.2 3)为供试材料 ,探讨了在充分供水-水分胁迫 -复水的处理过程中 ,短期不同遮光水平对棉花光合特性及其气孔响应的影响。结果表明 ,在水分处理过程中 ,所有不同遮光水平的棉花叶片对短期遮光具有相似的基本响应规律 :短期遮光使净光合速率迅速降低 ,气孔导度减少 ,但减少速率缓慢 ;遮阳网去掉后 ,叶片气孔重新开放速率和光合恢复被延迟。水分胁迫期间 ,所有遮光处理的水分利用效率均最高 ,但净光合速率均较低 ,光合及其气孔响应受到严重限制 ,遮光 75%受到的影响大于遮光 40 %的处理。复水后水分胁迫带来的残留影响仍然存在 ,结果使水分利用效率在整个水分处理过程中最低 ,遮光 75%的处理尤为明显。上述表明 ,土壤水分与短期遮光对棉花光合及其气孔响应的影响十分显著
Resumo:
利用光线的反射与聚焦成象原理 ,将地表高程转换成不同物象点位置的电信号 ,再经计算机处理成数字高程模型 ,即形成用于获取地表糙度数字高程模型的激光微地貌扫描仪。实验测量得到 :在相同初始条件下 ,不同坡度土槽放水和降雨实验后地表形态的数字地形模型。通过对侵蚀地表形态的分析 ,得出坡度对坡面侵蚀及沟蚀影响的某些初步规律。