782 resultados para indirect benefits


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Female mate choice influences the maintenance of genetic variation by altering the mating success of males with different genotypes. The evolution of preferences themselves, on the other hand, depends on genetic variation present in the population. Few models have tracked this feedback between a choice gene and its effects on genetic variation, in particular when genes that determine offspring viability and attractiveness have dominance effects. Here we build a population genetic model that allows comparing the evolution of various choice rules in a single framework. We first consider preferences for good genes and show that focused preferences for homozygotes evolve more easily than broad preferences, which allow heterozygous males high mating success too. This occurs despite better maintenance of genetic diversity in the latter scenario, and we discuss why empirical findings of superior mating success of heterozygous males consequently do not immediately lead to a better understanding of the lek paradox. Our results thus suggest that the mechanisms that help maintain genetic diversity also have a flipside of making female choice an inaccurate means of producing the desired kind of offspring. We then consider preferences for heterozygosity per se, and show that these evolve only under very special conditions. Choice for compatible genotypes can evolve but its selective advantage diminishes quickly due to frequency-dependent selection. Finally, we show that our model reproduces earlier results on selfing, when the female choice strategy produces assortative mating. Overall, our model indicates that various forms of heterozygote-favouring (or variable) female choice pose a problem for the theory of sexual ornamentation based on indirect benefits, rather than a solution.

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Paineilmaa käytetään teollisuudessa hyvin yleisesti. Teollistuneissa maissa teollisuuden käyttämästä energiasta 8-10 % kuluu paineilman tuottamiseen. Paineilmakompressoreiden nykyaikaisen ohjausjärjestelmän avulla voidaan säästää paineilmajärjestelmän energiakustannuksia jopa 30 %. Ohjauksen optimoinnin yhteydessä on havaittu epäkohtia joiden poistaminen on parantanut tuotantolaitosten toimintaan niin, että saadut säästöt ovat olleet jopa energiasäästöjä suuremmat. Näitä optimoinnin välillisiä hyötyjä haluttiin selvittää tarkemmin. Tutkimuksessa perehdyttiin paineilman käytön ongelmiin eri tuotantolaitoksissa ja arvioitiin paineilmajärjestelmän optimoinnin vaikutuksia tehtaiden tuotantoon. Koska paineilmaa käytetään monissa tehtaiden kriittisissä kohteissa, on tutkimuskohteiden paineilmajärjestelmän ongelmista aiheutunut merkittäviä tuotannonmenetyksiä ja lisäkustannuksia. Merkittävimmät paperitehtaiden tuotantoa vaikeuttavat paineilmajärjestelmän tekniset ongelmat liittyvät paineilman puhtausvaatimuksiin ja riittävän verkkopaineen ylläpidon vaikeuteen. Organisatorisia ongelmia ovat paineilmajärjestelmän käyttöhenkilöstön asiantuntemuksen puutteellisuudet. Näistä aiheutuvaa tuotannon epäluotettavuutta voidaan merkittävästi pienentää kehittämällä paineilmakompressoreita, jälkikäsittelyä, ohjausta ja valvontaa alan asiantuntijoiden avustuksella.

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Why females of many species mate multiply in the absence of direct benefits remains an open question in evolutionary ecology. Interacting and mating with multiple males can be costly to females in terms of time, resources, predation risk, and disease transmission. A number of indirect genetic benefits have been proposed to explain such behaviors, but the relative importance of these mechanisms in natural systems remains unclear. We tested for several direct and indirect benefits of polyandry in the walking stick Timema cristinae. We found no evidence of direct benefits with respect to longevity or fecundity. However, male x female genotypic interactions affected egg-hatching success and offspring production independent of relatedness, suggesting that mating with certain males benefits females and that the best male may differ for each female. Furthermore, multiply mated females biased paternity toward one or few males, and the extent of this bias was positively correlated to egg-hatching success. Our data, therefore, provide evidence for indirect benefits through compatibility effects in this species. By mating multiply, females may improve their chances of mating with a compatible male if compatibility cannot be assessed before mating. Such compatibility effects can explain the evolution and maintenance of polyandry in Timema and many other species.

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Male mating behaviors harmful to females have been described in a wide range of species. However, the direct and indirect fitness consequences of harmful male behaviors have been rarely quantified for females and their offspring, especially for long-lived organisms under natural conditions. Here, lifetime and intergenerational consequences of harmful male interactions were investigated in female common lizards (Lacerta vivipara) using field experiments. We exposed females to male harm by changing the population sex ratio from a normal female-biased to an experimental male-biased sex ratio during the first experimental year. Thereafter, females and their first generation of offspring were monitored during two additional years in a common garden with a female-biased sex ratio. We found strong immediate fitness costs and lower lifetime reproductive success in females subjected to increased male exposure. The immediate fitness costs were partly mitigated by direct compensatory responses after exposure to male excess, but not by indirect benefits through offspring growth, offspring survival, or mating success of offspring. These results support recent empirical findings showing that the direct costs of mating are not outweighed by indirect benefits.

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Yhteistyö yli organisaatiorajojen, huolellinen lähtötietojen selvittäminen, täsmällinen tehtävien suorittaminen sekä organisaatiossa olevan tiedon laaja hyödyntäminen ovat projektin onnistumisen kulmakiviä. Usein kuitenkin projekti hautautuu omaan arkeensa keskittyen liikaa tehtävien suorittamiseen, eikä luo katseita ympäristöönsä. Tässä työssä tavoitteena on ollut projekti- ja päätöksentekomallin luominen, joka ohjaa projektia heti alkumetreistä jo ennen kuin päätöstä sen toteuttamisesta on tehty. Tarkoituksena on myös ohjata yhteistyöhön yli organisaatiorajojen ja kaikkien näkökulmien huomioonottamiseen ennen projektin aloituspäätöstä. Malli luotiin konstruktiivisena tapaustutkimuksena ja se nojaa kirjallisuuteen melko laaja-alaisesti. Koska malli tulee ohjaamaan asiakasräätälöintejä toteuttavan organisaation projekteja, viitekehystä rajaavat räätälöinti, tuotehallinta ja ennen kaikkia projektin johtaminen. Tutkimusta varten on tutustuttu kohdeyrityksen prosesseihin ja liitetty konstruktio osaksi näitä prosesseja. Teorian pohjalle luotu konstruktio ratkaisee tutkimusongelman määrittämällä toimintamallin räätälöintiprojektien arvonmääritykseen ja toteutukseen. Se tuo järjestelmällisyyttä projektiehdokkaiden arvon määrittämiseen sekä johdonmukaistaa päätöksentekoprosessia ja projektin toteutusta ottaen huomioon suorien hyötyjen lisäksi epäsuorat hyödyt ja vaikutukset tuotetarjoomaan.

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The major objective of this chapter was to estimate the indirect benefits provided by the Cochin wetlands to direct, indirect and non-user populations.This chapter gives the details of the Contingent valuation survey that was executed in the study area. Section one described the actual survey and its execution. Section two undertook a detailed discussion of the methodological issues involved in the survey. Section three contained some discussion on the study.This analysis has demonstrated the feasibility of extending the use of contingent valuation methods to local populations in developing countries like India. Certain issues emerge from these applications. Income is strongly related to willingness to pay in these surveys, yet income levels are often low.Secondly, education is not a factor that influences willingness to pay in the coastal belt very much. Rather, relation of individual occupation to any wetland based activity very much influenced their willingness to pay. The study revealed that people very much valued the indirect function performed by wetlands, in fact as much as they valued the direct benefits provided by the system. There still exist differences of opinions among experts when undertaking such valuation studies. However, in the absence of a better technique for valuing environmental services that have no markets, this is definitely a first step

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o trabalho apresenta a análise de um programa realizado pela Prefeitura do Município de Piraí, localizado no Estado do Rio de Janeiro, no qual visa estruturar uma política de geração de empregos a partir da cadeia produtiva de determinados produtos. A execução do programa teve início, com o planejamento e a construção de um condomínio industrial, que pudesse não só gerar empregos nas indústrias nele instaladas, mas também proporcionasse oportunidades de geração de empregos, nos estágios anteriores e/ou posteriores desta possível cadeia produtiva gerada. A participação do município então se estenderia à capacitação da população, para que esta, organizada principalmente com a estruturação de cooperativas, com apoio financeiro municipal, pudesse agregar-se aos estágios da produção. O enfoque municipal também se estendeu à forma de trabalho que demonstrasse potencialidade na região, sendo o caso do incentivo à piscicultura. Com isso, a geração de empregos se baseou em três grandes áreas: na indústria, na cooperativa e na piscicultura. Este programa proporcionou beneficios indiretos, além dos diretamente verificados, dados os objetivos preliminares, como uma conseqüente melhoria no nível de renda da população e um crescimento ordenado da região. A gestão pública enfatizou a necessidade de estabelecer uma forma de comércio integrado dos canais de produção local, visando proporcionar um nível de demanda satisfatório à manutenção dos mesmos. O beneficio criado pelo governo do município de Piraí, além da melhoria nos índices tradicionais referentes à qualidade de vida, pode ser observado numa maior interação do processo de cidadania, por meio das entidades comunitárias, associações e cooperativas.

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The urban growth without the prior infrastructure has caused many environmental impacts such as the damage to quality of the water resources in the cities. Along with natural scarcity in some regions, this is one of the factors that limit the availability of drinking water. As a result, the conservation of drinking water is becoming one of the major concerns in sustainable architectural projects. Within this context, this dissertation proposes to develop the design of an educational building focusing on water consumption rationalization. The proposed project is located in UFRN Campus at Currais Novos, an area of warm and dry climate and low rainfall. The proposal seeks to integrate ways to reduce water consumption o to architecture, in order to exploit the advantages and savings. After quantifying the benefits achieved, it was concluded that it is possible to reduce significantly the drinking water consumption in educational buildings in universities using three principles: reduction the water consumption at the point of use, replacement of the water source and internal recycling. Calculations and simulations indicated that the proposed building may have water consumption up to 56% lower than if it would be provided by conventional facilities. Rationalization of water consumption brings direct and indirect benefits, with influences on the environmental, social and economic fields

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One of the ways of Geography is the study of the relationship between man and space, so that the question of the preservation of the environment and yours attributes is increasingly standing out in society. The State Forest Edmundo Navarro de Andrade is a conservation of sustainable use, so with this category aims to reconcile nature conservation with sustainable use of a portion of its resources, and a place that has many direct and indirect benefits to the interconnected environment . Therefore, the object of study is the State Forest Edmundo Navarro de Andrade, highlighting its Public Use Area. This work has as main objective to measure the importance of the State Forest Edmundo Navarro de Andrade, in the view of visitors, and present some key aspects of the unit in relation to its attractions. Thus, we seek the appreciation of the history of the city of Rio Claro and qualities of the State Forest Edmundo Navarro de Andrade

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Mounting an immune response against pathogens incurs costs to organisms by its effects on important life-history traits, such as reproductive investment and survival. As shown recently, immune activation produces large amounts of reactive species and is suggested to induce oxidative stress. Sperm are highly susceptible to oxidative stress, which can negatively impact sperm function and ultimately male fertilizing efficiency. Here we address the question as to whether mounting an immune response affects sperm quality through the damaging effects of oxidative stress. It has been demonstrated recently in birds that carotenoid-based ornaments can be reliable signals of a male's ability to protect sperm from oxidative damage. In a full-factorial design, we immune-challenged great tit males while simultaneously increasing their vitamin E availability, and assessed the effect on sperm quality and oxidative damage. We conducted this experiment in a natural population and tested the males' response to the experimental treatment in relation to their carotenoid-based breast coloration, a condition-dependent trait. Immune activation induced a steeper decline in sperm swimming velocity, thus highlighting the potential costs of an induced immune response on sperm competitive ability and fertilizing efficiency. We found sperm oxidative damage to be negatively correlated with sperm swimming velocity. However, blood resistance to a free-radical attack (a measure of somatic antioxidant capacity) as well as plasma and sperm levels of oxidative damage (lipid peroxidation) remained unaffected, thus suggesting that the observed effect did not arise through oxidative stress. Towards the end of their breeding cycle, swimming velocity of sperm of more intensely colored males was higher, which has important implications for the evolution of mate choice and multiple mating in females because females may accrue both direct and indirect benefits by mating with males having better quality sperm.

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Background. Obesity in America has increased exponentially since the 1970s with no sign of slowing down. It is a major public health problem, and is currently the second leading cause of preventable deaths in America (Flegal et al., 2010). Bariatric surgery is currently the only approved therapy that has shown to have a lasting impact on obese patients. While the initial cost of the surgery remains high, numerous cost-benefit analyses have demonstrated an overall cost saving within two to five years (McEwen et al., 2010). Only three states, including Texas, do not currently fund bariatric surgery through Medicare and Medicaid. ^ Objectives. To determine whether the current data on the cost-benefit analysis of bariatric surgery supports Texas' decision to not publicly fund bariatric surgery through its Medicare and Medicaid programs. ^ Methods. We conducted literature reviews to determine the current cost of obesity in Texas as well as the methods being employed to treat obesity currently. We then analyzed the history of bariatric surgery and its current implementation, looking at safety and the future benefits of bariatric surgery. We then looked at key cost-benefit analyses and meta-analyses to determine the cost effectiveness of bariatric surgery. We then analyzed both direct medical expenditures and indirect benefits of bariatric surgery. ^ Conclusions. If the obesity epidemic continues unabated, it will become one of the leading health expenditures in Texas within decades. Given that surgery is currently the only approved therapy for obesity that has been shown to be effective in the majority of patients, Texas' decision not to publicly fund bariatric surgery is short sighted.^

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Se determinó la factibilidad económica de la actividad conjunta de recría y engorde a corral de bovinos para carne en la provincia de Mendoza, postulando la hipótesis que la rentabilidad del engorde a corral puede mejorar al integrarse con la recría. Las alternativas analizadas fueron: a) riego superficial y confección de rollos de alfalfa con contratista (alternativa 1.1) o con maquinaria propia (alternativa 1.2) y b) riego por aspersión con cañón regador y confección de rollos de alfalfa con contratista (alternativa 2.1) o con maquinaria propia (alternativa 2.2). Los precios usados para valorizar inversiones, gastos operativos e ingresos fueron los vigentes en abril de 2011. Fue determinada la cantidad de animales por año necesarios para alcanzar rentabilidad en dichas alternativas, con y sin la percepción de la compensación provincial. Se estimó la tasa interna de retorno, considerándose una tasa de interés anual del 12% como costo de oportunidad del capital. La rentabilidad del engorde a corral mejoró al integrarse con la recría. La unidad económica en la actividad conjunta varió entre 446 (alternativa 1.1, con compensación) y 708 animales por año (alternativa 2.2, sin compensación). Los equipos para confección de rollos de alfalfa y riego proporcionan beneficios directos: autonomía en la confección de rollos y mayor eficiencia de riego, e indirectos: ingresos adicionales por la prestación del servicio de confección de rollos a terceros.

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The bed bug, Cimex lectularius, has a unique mode of copulation termed “traumatic” insemination [Carayon, J. (1966) in Monograph of the Cimicidae, ed. Usinger, R. (Entomol. Soc. Am., Philadelphia), pp. 81–167] during which the male pierces the female's abdominal wall with his external genitalia and inseminates into her body cavity [Carayon, J. (1966) in Monograph of the Cimicidae, ed. Usinger, R. (Entomol. Soc. Am., Philadelphia), pp. 81–167]. Under controlled natural conditions, traumatic insemination was frequent and temporally restricted. We show for the first time, to our knowledge, that traumatic insemination results in (i) last-male sperm precedence, (ii) suboptimal remating frequencies for the maintenance of female fertility, and (iii) reduced longevity and reproductive success in females. Experimental females did not receive indirect benefits from multiple mating. We conclude that traumatic insemination is probably a coercive male copulatory strategy that results in a sexual conflict of interests.

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The net effect of sexual selection on nonsexual fitness is controversial. On one side, elaborate display traits and preferences for them can be costly, reducing the nonsexual fitness of individuals possessing them, as well as their offspring, In contrast, sexual selection may reinforce nonsexual fitness if an individual's attractiveness and quality are genetically correlated. According to recent models, such good-genes mate choice should increase both the extent and rate of adaptation. We evolved 12 replicate populations of Drosophila serrata in a powerful two-way factorial experimental design to test the separate and combined contributions of natural and sexual selection to adaptation to a novel larval food resource. Populations evolving in the presence of natural selection had significantly higher mean nonsexual fitness when measured over three generations (13-15) during the course of experimental evolution (16-23% increase). The effect of natural selection was even more substantial when measured in a standardized, monogamous mating environment at the end of the experiment (generation 16; 52% increase). In contrast, and despite strong sexual selection on display traits, there was no evidence from any of the four replicate fitness measures that sexual selection promoted adaptation. In addition, a comparison of fitness measures conducted under different mating environments demonstrated a significant direct cost of sexual selection to females, likely arising from some form of male-induced harm. Indirect benefits of sexual selection in promoting adaptation to this novel resource environment therefore appear to be absent in this species, despite prior evidence suggesting the operation of good-genes mate choice in their ancestral environment. How novel environments affect the operation of good-genes mate choice is a fundamental question for future sexual selection research.