East Gondwana ancestry of the sunflower alliance of families

The sunflower alliance of families comprises nearly 10% of all flowering plant species and includes the largest of all plant families, the sunflower family Asteraceae, which has 23,000 species, and the bellflower family Campanulaceae. Both are worldwide in distribution, but the majority of their species occur in the northern hemisphere. Recently it has been shown that a number of small, woody families from the Australian–Southwest Pacific area also belong in this relationship. Here we add yet another such family and present phylogenetic, biogeographic, and chronological analyses elucidating the origin of this large group of plants. We show that the ancestral lineages are confined to Malesia, Australia, New Guinea, and New Zealand and that the sunflower and bellflower families represent phylogenetically derived lineages within a larger group with a Cretaceous and southern-hemisphere, presumably East Gondwana, ancestry. Their highly derived position in the flowering plant phylogeny makes this significant for understanding the evolution of flowering plants in general.

Los Angeles, California (East), and vicinity, 1953 (Raster Image) (Image 1 of 2)

This layer is a georeferenced raster image of the United States Geological Survey sheep map set entitled: Los Angeles and vicinity, East [and West], California. Edition 1953. It was published in 1956. Compiled from 1:24,000 scale maps of the Azusa 1953, Mt. Wilson 1953, Pasadena 1953, Los Angeles 1953, El Monte 1953, Baldwin Park 1953, La Habra 1950, Whittier 1949, and South Gate 1952 7.5 minute quadrangles. Scale 1:24,000. This layer is image 1 of 2 total images of the two sheet source map set representing the eastern portion of the map set. The image inside the map neatline is georeferenced to the surface of the earth and fit to the California State Plane Zone V Coordinate System NAD27 (in Feet) (Fipszone 0405). All map collar and inset information is also available as part of the raster image, including any inset maps, profiles, statistical tables, directories, text, illustrations, index maps, legends, or other information associated with the principal map. USGS maps are typical topographic maps portraying both natural and manmade features. They show and name works of nature, such as mountains, valleys, lakes, rivers, vegetation, etc. They also identify the principal works of humans, such as roads, railroads, boundaries, transmission lines, major buildings, etc. Relief is shown with standard contour intervals of 5 and 25 feet. Please pay close attention to map collar information on projections, spheroid, sources, dates, and keys to grid numbering and other numbers which appear inside the neatline. This layer is part of a selection of digitally scanned and georeferenced historic maps from The Harvard Map Collection as part of the Imaging the Urban Environment project. Maps selected for this project represent major urban areas and cities of the world, at various time periods. These maps typically portray both natural and manmade features at a large scale. The selection represents a range of regions, originators, ground condition dates, scales, and purposes.