201 resultados para palaeontology


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Palaeogeographic and tectono-sedimentary interpretation of northern Portugal, in which previous studies (geomorphology, lithostratigraphy, mineralogy, sedimentology, palaeontology, etc.) were considered, is here proposed. Cenozoic shows different features according to its morphotectonic setting in the eestern region (Trás-os-Montes) or near to the Atlantic coast (western region, Minho and Douro Litoral areas). Although in the eastern region the sedimentary record is considered late Neogene, in some places Paleogene (?) was identified. This oldest record, represented by alluvial deposits, was preserved from complete erosion because of its position inside Bragança-Vilariça-Manteigas fault zone grabens. Later sedimentary episodes (upper Tortonian-Zanclean ?), represented by two allostratigraphical units, were interpreted as proximal fluvial braided systems of an endorheic hydrographic network, draining to the Spanish Duero Basin (eastwards); nowadays, they still remained in tectonic depressions and incised-valleys. Later on, eastern sedimentation becomes scarcer because Atlantic fluvial systems (e.g. the pre-Douro), successively, captured previous endorheic drainages. The proximal reaches of the allostratigraphic unit considered Placencian is recorded in Mirandela (western Trás-os-Montes) but the following fluvial episode (Gelasian-early Pleistocene ?) was already documented in east Trás-os-Montes, preserved in high platforms and in tectonic depressions. Placencian and Quaternary sedimentary records in the western coastal zone, mainly represented by terraces, are located in the Minho, Lima, Alverães, Cávado and Ave large fluvial valleys and in the Oporto littoral platform. In conclusion, northern Portugal Tertiary sedimentary episodes were mainly controlled by tectonics, but later on (Placencian-Quaternary) also by eustasy.

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Des dinosaures sur papier : des notes « sur le terrain » comme on en trouve dans le Badlands de Robert Kroetsch passe en revue cette œuvre postmoderne de 1975 portant sur une expédition paléontologique fictive près de la rivière Red Deer, en Alberta, conformément à la récente tendance à exiger la vérification systématique des données à la base des récits métafictifs historiographiques dans la littérature canadienne-anglaise. Inspirée de l’exploration canonique qu’a effectuée John Livingston-Lowes des plus grands poèmes de Samuel Taylor Coleridge par le biais de la mine d’or du Gutch Memorandum Book dans The Road to Xanadu, cette thèse entreprend un nouveau type de recherche qui se démarque des archives conventionnelles et de la tradition documentaire. S’appuyant sur des documents holographes non publiés provenant de dépôts d’archives situés au Québec, en Ontario et en Alberta et écrits par des collecteurs, des géologues et des paléontologues de la Commission géologique du Canada, ainsi que sur des notes « sur le terrain », des notes de recherche et des journaux personnels écrits par Robert Kroetsch pendant la rédaction de son roman Badlands, cet examen critique révèle les strates sous-jacentes inédites d’une œuvre de fiction particulière. Dans pratiquement toute fouille paléontologique, le retrait de ce qui enveloppe un spécimen révèle souvent des données supplémentaires qui peuvent, si elles sont soigneusement interprétées, offrir des indices essentiels sur les environnements paléontologiques. Ainsi, un squelette de dinosaure est rarement retiré d’une carrière stérile dans son intégralité. Il en va de même pour toute recherche sur un processus littéraire. Aucun texte ne s’autosuffit. Comme Kroetsch s’est efforcé de produire son récit sous forme d’interrogation sur la création et la transmission des données historiques, particulièrement grâce à des notes « sur le terrain », une vaste étude de ce « terrain » comprenant des intertextes de l’Antiquité, des sciences, de l’Histoire, de l’histoire populaire, de récits de voyages et de la littérature canadienne et internationale est ici menée. On y fait librement référence à des périodes et à des auteurs très diversifiés, allant de Thomas Jefferson et des tombelles à Bruce Chatwin et sa peau de « brontosaure ». Évidemment, aucune entreprise interdisciplinaire du genre ne peut être exhaustive. Ce projet se veut plutôt une vitrine littéraire réunissant des curiosités autour d’une œuvre principale, soit le Badlands de Robert Kroetsch. Réduites à leur plus simple expression, les notes « sur le terrain » constituent des messages destinés à la postérité. En explorant trois thèmes principaux, cette thèse explique comment ces messages pourraient être transmis. « Saxa Loquuntur ! », ainsi intitulé en référence à l’analogie de Freud avec l’archéologie, traite des métaphores associées aux témoignages de la pierre ; « Good Jones » porte sur les façons dont la taxinomie peut combler le désir d’un chercheur d’os de ne pas tomber dans l’oubli ; « Box 16 » suit une piste documentaire en parcourant les écrits de Kroetsch pour reconstituer tant l’élaboration d’un roman que les notions de temps, d’espace et d’origine d’une œuvre littéraire.

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La version intégrale de cette thèse est disponible uniquement pour consultation individuelle à la Bibliothèque de musique de l’Université de Montréal (www.bib.umontreal.ca/MU).

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Phosphorite-filled crustacean burrows associated with a Campanian-age omission surface in the north-western Negev are described. The phosphatic burrow casts weather out displaying scratches (bioglyphs) and two types of local swellings (chambers), which are flattened normal to the course of the burrow. The more abundant chamber type is a flattened spheroid (diameter 45-50 mm) or a flattened, highly prolate ellipsoid of larger dimensions, with bioglyphs. The other type is a flattened spheroid (diameter 45 mm), gently rounded on the upper side and flat on the base. Rings of elevations on the cast (representing moats) form interconnected circlets, each capped by about eight rounded hemispherical tubercles (4 x 4 mm) (pits on original), the whole forming a discrete network. The first type of chamber may have hosted the young (nursery chamber) and/or stored food. The second type of cast replicates a chamber with a pitted floor, which may have formed a brood chamber for 60-70 spherical eggs, each about 3 mm in diameter. Brood chambers in crustacean burrow systems were previously suspected, but only at burrow terminations. The interpreted K-type breeding strategy, brood care and associated functions require a high degree of social organization, none of which has been observed in extant crustaceans, but all occur within social insects.

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Results of a cladistic analysis of the suborder Conulariina Miller and Gurley, 1896, a major extinct (Vendian-Triassic) group of scyphozoan cnidarians, are presented. The analysis sought to test whether the three conulariid subfamilies (Conulariinae Walcott, 1886, Paraconulariinae Sinclair, 1952 and Ctenoconulariinae Sinclair, 1952) recognized in the Treatise on Invertebrate Paleontology ( TIP) are monophyletic. A total of 17 morphological characters were scored for 16 ingroup taxa, namely the genera Archaeoconularia, Baccaconularia, Climacoconus, Conularia, Conulariella, Conularina, Ctenoconularia, Eoconularia, Glyptoconularia, Metaconularia, Notoconularia, Paraconularia, Pseudoconularia, Reticulaconularia, Teresconularia and Vendoconularia. The extant medusozoan taxa Cubozoa, Stauromedusae, Coronatae and Semaeostomeae served as outgroups. Unweighted analysisof the data matrix yielded 1057 trees, and successive weighting analysis resulted in one of the 1057 original trees. The ingroup is monophyletic with two autapomorphies: (1) the quadrate geometry of the oral region; and (2) the presence of a mineralized (phosphatic) periderm. Within the ingroup, the clade (Vendoconularia, Teresconularia, Conularina, Eoconularia) is supported by the sinusoidal longitudinal geometry of the transverse ridges, and the much larger clade (Baccaconularia, Glyptoconularia, Metaconularia, Pseudoconularia, Conularia, Ctenoconularia, Archaeoconularia, Notoconularia, Climacoconus, Paraconularia, Reticulaconularia) is supported by the presence of external tubercles, which, however, were lost in the clade (Notoconularia, Climacoconus, Paraconularia, Reticulaconularia). As proposed by Van Iten et al. (2000), the clade (Notoconularia, Climacoconus, Paraconularia, Reticulaconularia) is supported by the termination and alternation of the transverse ribs in the corner sulcus. The previously recognized subfamilies Conulariinae, Paraconulariinae and Ctenoconulariinae were not recovered from this analysis. The diagnostic features of Conulariinae (continuation of the transverse ornament across the corner sulcus and lack of carinae) and Ctenoconulariinae ( presence of carinae) are symplesiomorphic or homoplastic, and Paraconulariinae is polyphyletic. The families Conulariellidae Kiderlen, 1937 and Conulariopsidae Sugiyama, 1942, also recognized in the TIP, are monogeneric, and since they provide no additional phylogenetic information, should be abandoned.

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Sir Frederick McCoy, during a long career involved with taxonomy, contributed extensively to the knowledge of the fossil record of the Phylum Brachiopoda, From his classic early monographs on the fossil faunas of the Carboniferous and Silurian of Ireland, to his later works in Victoria where important new species were described and illustrated, McCoy demonstrated the same care, meticulous rigour and quality of illustrations that typified all his work. His contributions on the Brachiopoda are of high and long-lasting significance but form only part of his much broader contribution to palaeontology.

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Frederick McCoy contributed to the knowledge of the fossil record of the Tasmanian Devil Sarcophilus harrisii Boitard, 1842 in Victoria by including a number of figured specimens in the Prodromus of the Palaeontology of Victoria. However, an article McCoy wrote under the pseudonym 'Microzooni highlighted his anti-Darwinian thoughts and embraced a successionist viewpoint. The article, entitled 'Pre-historic Tasmanian Devils', is an interesting account of zoogeography from a successionist perspective, and is used here to contrast McCoy's anti-evolutionary viewpoint with modern Darwinian thought. A number of fossil sarcophilines discovered since McCoy's death illustrate the shortcomings of McCoy's favoured anti-Darwinian viewpoint when discussing the nature of evolution and extinction.

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From August 1869 until May 1871, an anonymous naturalist under the pseudonym 'Microzoon' published a superb series of articles in a weekly Melbourne newspaper, The Australasian. The author was undoubtedly Frederick McCoy. The Microzoon articles provide a valuable early record of
aspects of the natural history of Victoria, in particular the bird life, but also covering a selection of other topics including snakes, insects, fish, molluscs, geology, palaeontology and stratigraphy.

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The terms ‘Leptodus Shales’ and ‘Leptodus Beds’ have been used to describe a rich brachiopod bearing unit within the Permian argillaceous facies of the Central Belt of Peninsular Malaysia. To date there has been no formal description of this unit regarding its age, spatial distribution or faunal composition. A review of previous literature, backed by recently collected data from our field surveys and biostratigraphical studies reveals that there is a sequence of fossiliferous assemblages within the Leptodus Shales, which range in age from Middle Permian to possibly early Late Permian and extend geographically from southern Kelantan to southern Pahang, Peninsular Malaysia. These assemblages are found in argillaceous sediments which are often highly tuffaceous, and in northern Pahang are associated with pyroclastic volcanics of probable island-arc origin. The faunas are of Palaeo-equatorial affinity and are taxonomically close to faunas in Indochina, such as the Sisophon fauna in Cambodia. Typical elements include Vediproductus cf. punctatiformis (Chao), Transennatia gratiosa (Waagen), T. termierorum Sone, Leman and Shi, Uncuninellina timorensis (Beyrich), Leptodus richthofeni Kayser, L. cf. tenuis (Waagen), Leptodus nobilis (Waagen), Gubleria aff. ninglangensis Fang and Jiang, and Spyridiophora gubleri Termier and Termier.

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The brachiopod fauna from the Tupe Formation at La Herradura Creek, located on the west flank of Perico Hill, San Juan Province, Argentina, palaeogeographically belongs to the western sector of the Paganzo basin ('Guandacol embayment'). The stratigraphical section of the Tupe Formation at La Herradura Creek is the stratotype of the Tivertonia jachalensis-Streptorhynchus inaequiornatus biozone, was previously regarded as being of Late Carboniferous age but here is assigned to the earliest Permian (Asselian). We describe and review the biozone assemblage, which consists of Streptorhynchus inaequiornatus, Tivertonia jachalensis, Kochiproductus sp., Costatumulus sp., Coronalosia argentinensis, Tupelosia paganzoensis, Trigonotreta pericoensis, Septosyringothyris sp. aff. Septosyringothyris jaguelensis and Crurithyris? sp. This brachiopod assemblage is related to Indian and Australian Early Permian faunas and its presence in the La Herradura Creek section provides new evidence in support of an Asselian (Early Permian) age for the Tivertonia jachalensis-Streptorhynchus inaequiornatus biozone. This assemblage is also important for intra- and inter-basinal correlation because several of its characteristic species have been identified from other sections of the Paganzo basin and the Riacuteo Blanco basin. The proposed age for this biozone is consistent with the age of palynological data from slightly above the marine faunas from the stratotype locality.

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The Permian-Triassic extinction pattern in the peri-Gondwanan region is documented biostratigraphically, geochemically and sedimentologically based on three marine sequences deposited in southern Tibet and comparisons with the sections in the Salt Range, Pakistan and Kashmir. Results of biostratigraphical ranges for the marine faunas reveal an end-Permian event comparable in timing with that known at the Meishan section in low palaeolatitude as well as Spitsbergen and East Greenland in northern Boreal settings although biotic patterns earlier in the Permian vary. The previously interpreted delayed extinction (Late Griesbachian) at the Selong Xishan section is not supported by our analysis. The end-Permian event exhibits an abrupt marine faunal shift slightly beneath the Permian-Triassic boundary (PTB) from benthic taxa- to nektic taxa-dominated communities. The climate along the continental margin of Neo-Tethys was cold before the extinction event. However, a rapid climatic warming event as indicated by the southward invasion of abundant warm-water conodonts, warm-water brachiopods, calcareous sponges, and gastropods was associated with the extinction event. Stable isotopic values of δ13Ccarb, δ13Corg and δ18O show a sharp negative drop slightly before and during the extinction interval. Sedimentological and microstratigraphical analysis reveals a Late Permian regression, as marked by a Caliche Bed at the Selong Xishan section and the micaceous siltstone in the topmost part of the Qubuerga Formation at the Qubu and Tulong sections. The regression was immediately followed by a rapid transgression beneath the PTB. The basal Triassic rocks fine upward, and are dominated by dolomitic packstone/wackestone containing pyritic cubes, bioturbation and numerous tiny foraminifers, suggesting that the studied sections were deposited during the initial stage of the transgression and hence may not have been deeply affected by the anoxic event that is widely believed to characterise the zenith of the transgression.

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This thesis deals with the stratigraphy and brachiopod systematic palaeontology of the latest Devonian (Famennian) to Early Permian (Kungurian) sedimentary sequences of the Tarim Basin, NW China. Brachiopod faunas of latest Devonian and Carboniferous age have been published or currently in press in the course of the Ph.D candidature and are herein appendixed, while the Early Permian brachiopod faunas are systematically described in this thesis. The described Early Permian brachiopod faunas include 127 species, of which 29 are new and 12 indeterminate, and six new genera (subgenera) are proposed; Tarimella, Bmntonella, Marginifera (Arenaria), Marginifera (Nesiotia), Baliqliqia and Ustritskia. A new integrated brachiopod biostratigraphical zonation scheme is proposed, for the first time, for the latest Devonian-Early Permian sequences of the entire Tarim Basin on the basis of this study as well as previously published information (including the Candidate's own published papers). The scheme consists of twenty three brachiopod acm biozones, most of which replace previously proposed assemblage or assemblage zones. The age and distribution of these brachiopod zones within the Tarim Basin and their relationships with other important fossil groups are discussed. In terms of regional correlations and biostratigraphical affinities, the Late Devonian to Early Carboniferous brachiopod faunas of the Tarim Basin are closest to those from South China, while the Late Carboniferous faunas demonstrate strong similarities to coeval faunas from the Urals, central Asia, North China and South China. During the Asselian-Sakmarian, strong faunal links between the Tarim Basin and those of the Urals persisted, while at the same time links with central Asia, North China and South China weakened. On the other hand, during the Artinskian-Kungurian times, affinities of the Tarim faunas with the Urals/Russian Platform rapidly reduced, when those with peri-Gondwana (South Thailand, northern Tibet) and South China increased. Thirty lithofacies (or microfacies) types of four facies associations are recognised for the Late Devonian to early Permian sediments. Based on detailed lithostratigraphy, biostratigraphy and facies analysis, 23 third-order sequences belonging to four supcrsequences are identified for the Late Devonian to Early Permian successions, from which sea-level fluctuation curves are reconstructed. The sequence stratigraphical analysis reveals that four major regional regressions, each marking a distinct supersequence boundary, can be recognised; they correspond to the end-Serpukhovian, end-Moscovian, late Artinskian and end-Kungurian times, respectively. The development of these sequences is considered to have been formed and regulated by the interplay of both eustasy and tectonism. Using the system tract of a sequence as the mapping time unit, a succession of 47 palaeogeographical maps have been reconstructed through the Late Devonian to Early Permian. These maps reveal that the Tarim Basin was first immersed by southwest-directed (Recent geographical orientation) transgression in the late Famennian after the Caledonian Orogeny. Since then, the basin had maintained its geometry as a large, southwest-mouthed embayment until the late Moscovian when most areas were the uplifted above sea-level. The basin was flooded again in late Asselian-Artinskian times when a new transgression came from a large epicontinental sea lying to its northwest. Thereafter, marine deposition was restricted to local areas (southwestern and northwestern margins until the late Kungurian, while deposition of continental deposits prevailed and continued through the Middle and late Permian into the Triassic.

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A brachiopod fauna comprising nine species in eight genera from three closely spaced stratigraphic horizons of the same stratigraphic section is described for the first time from the Laibin Limestone in the uppermost part of the Maokou Formation in the Guadalupian/Lopingian (G/L) GSSP section at Penglaitan, Guangxi Autonomous Region, South China. The brachiopod assemblages are bracketed between two conodont zones: Jinogondolella xuanhanensis Zone below and Jinogondolella granti Zone above and, therefore, they can be safely assigned to the latest Capitanian in age. However, all but one of the nine brachiopod species from the Laibin Limestone carry strong early Lopingian (Wuchiapingian) aspect. Thus, the discovery of this brachiopod fauna not only suggests that some Lopingian brachiopod species had already appeared in the late Guadalupian (Capitanian); more importantly, it has also highlighted the fact that both the previously noted pre-Lopingian life crisis (or end-Guadalupian or Middle Permian mass extinction) and Lopingian recovery/radiation actually occurred in late Capitanian times, sometime before the G/L chronostratigraphic boundary. So far, the Penglaitan GSSP section provides the highest-resolution disappearance patterns of different fossil groups around the G/L boundary.

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This research systematically studied Permian brachiopod fossils from southeastern Mongolia and correlated them with many places around East Asia. The research found that the fossil faunas are on the mixed affinities between high-latitude cool- to cold-water and low-latitude warm water species. Explanations are provided through southeastern Mongolia’s unique palaeogeographical and palaeoceanographical position in the Permian time.