Linshuichonetes gen. nov., a new rugosochonetid (Brachiopoda) genus from the Liangshan Formation (Early Permian) in Sichuan, China, and its ecology

A new genus and species, Linshuichonetes elfinis, belonging to the Rugosochonetidae, is described from the Early Permian (Late Artinskian or early Kungurian) Liangshan Formation of the Yangtze block. The new genus is defined externally by the presence of fine, but delayed, capillation and a weak or absent median sulcus and fold and by the presence of a distinct posteromedian sinus on the ventral umbo; and internally by a lack of median, lateral and accessory septa in the dorsal interior; absence of vascular trunks in the ventral interior and the presence of distinct radiating rows of papillae in the interior of both valves, particularly an unusual clustered arrangement of papillae on the posteromedian portion of the dorsal interior. The local environment during the deposition of the Liangshan Formation appears to have been a restricted tidal flat or lagoon which experienced frequent sealevel fluctuations associated with the onset of the Yanghsingian transgression. The new species, L. elfinis, appears to have several morphological adaptations enabling successful exploitation of this environment. It was typically a very small and thin-valved species with a high surface area to volume ratio, an advantage in an oxygen restricted environment. The small size and numerous body spinules would have aided individuals to remain suspended at the top of the fine, soft substrate. It also dominated the brachiopod assemblage in the Liangshan Formation, comprising up to 94%of specimens within a bed. These factors indicate that the new species appears to be an opportunistic species.

Guadalupian (Middle Permian) brachiopods from Sungai Toh, a Leptodus Shale locality in the Central Belt of Peninsular Malaysia. Part I: Lower Horizons

This work presents a systematic study of Permian Brachiopoda from the Sungai Toh Leptodus Shale locality, Pahang State, Peninsular Malaysia. This locality lies within the Central Belt of Peninsular Malaysia, a tectonic unit characterised.by tuffaceous sediments and limestones of Late Palaeozoic age. Two brachiopod-bearing horizons were studied in detail at this locality, the lower one (Horizon 2) bearing a mixed plant and invertebrate assemblage, including the brachiopods Urushtenoidea chaoi (CHING), Leptodus richthofeni KAYSER, Anidanthus cf. sinosus HUANG, Acosarina dorashamensis (SOKOLSKAJA), A. minuta (ABleH) and unidentifiable species of Linoproduetus, Neochonetes, and Strophalosiina. Horizon 3 contains a more abundant and diverse brachiopod fauna, comprising a total. of 57 species representing 47 genera, including Vediproductus punetatiformis (CHAO), Permianella typica HE & ZHU, Tranrennatia gratiosa (WAAGEN), Leptodus richthofeni KAYSER, Leptodus cf. tenuis (WAAGEN) and "Semibrachythyrina" [= Alphaneospirifer] cf. pyramidiformis LIANG. It is
suggested in this study that the age of the Sungai Toh locality is Capitanian (late Guadalupian) to possibly Wuchiapingian (early Lopingian),
as it appears to correlate well with the Lengwu fauna from Zhejiang in eastern China. The palaeobiogeographical affinities of the Sungai Toh fauna are interesting, mainly indicating strong Palaeo-equatorial affinities, while there are also some elements more typical of the cooler periGondwana
Region.

Paleobiogeographical extinction patterns of Permian brachiopods in the Asian-western Pacific region

Spatial and temporal variations in biological diversity are critical in understanding the role of biogeographical regulation (if any) on mass extinctions. An analysis based on a latest database of the stratigraphic ranges of 89 Permian brachiopod families, 422 genera, and 2059 species within the Boreal, Paleoequatorial, and Gondwanan Realms in the Asian–western Pacific region suggests two discrete mass extinctions, each possibly with different causes. Using species/family rarefaction analysis, we constructed diversity curves for late Artinskian–Kungurian, Roadian–Wordian, Capitanian, and Wuchiapingian intervals for filtering out uneven sampling intensities. The end-Changhsingian (latest Permian) extinction eliminated 87–90% of genera and 94–96% of species of Brachiopoda. The timing of the end-Changhsingian extinction of brachiopods in the carbonate settings of South China and southern Tibet indicates that brachiopods suffered a rapid extinction within a short interval just below the Permian/Triassic boundary.

In comparison, the end-Guadalupian/late Guadalupian extinction is less profound and varies temporally in different realms. Brachiopods in the western Pacific sector of the Boreal Realm nearly disappeared by the end-Guadalupian but experienced a relatively long-term press extinction spanning the entire Guadalupian in the Gondwanan Realm. The end-Guadalupian brachiopod diversity fall is not well reflected at the timescale used here in the Paleoequatorial Realm because the life-depleted early Wuchiapingian was overlapped by a rapid radiation phase in the late Wuchiapingian. The Guadalupian fall appears to be related to the dramatic reduction of habitat area for the brachiopods, which itself is associated with the withdrawal of seawater from continental Pangea and the closure of the Sino-Mongolian seaway by the and-Guadalupian.

Permian soft-sediment deformation structures related to earthquake in the Southern Sydney Basin, Eastern Australia

Sydney Basin is located in the eastern part of Australia, Lachlan Fold Belt, and between the New England Fold Belt. From the Sydney basin at the end of the Late Carboniferous to Middle Triassic experienced back-arc spreading to the foreland basin at different stages: back-arc spreading stage (Carboniferous ), A passive thermal subsidence stage (early in the Permian Berry) and load deflection extruding stage (in Broughton Permian - Triassic). This time at the Sydney basin on the eastern side of the New England Fold Belt for the island Background of the arc. As a result, back-arc in the Permian Basin of the South Sydney basin by the back-arc spreading the eastern side of the arc and trench subduction before the impact of strong seismic activity, the development of a series of earthquake-related seismites to form various types and Seismic activity related to the deformation of soft sediment structure. Permian Basin, South Sydney's soft sediment deformation including cracks in shock-fold, liquefied vein, volcanic sand, load structure, flame Construction, pillow-like structure, spherical structure, pillow Layer structure slump, and so breccia. To which the cracks in shock-fold fibrillation is a direct result of earthquake faults and folds; pillow is a layer of sand caused by the earthquake fibrillation dehydration, the formation of the sinking; liquefied vein, Volcanic sand for the liquefaction of sand penetration of the formation of earthquake fissures formed; load structure, flame Construction, pillow-like structure, spherical structure is affected by the earthquake fibrillation in the sand, mudstone interface because of the sinking sand, mud layer formed through ; Slump structures and breccia of the earthquake was caused by the gravitational collapse or the formation of the debris flow. Fissures, earthquake-fold, liquefied vein, volcanic sand, load structure, flame Construction, pillow-like structure, spherical structure, pillow-like layer Equivalent to the original earthquake rocks the plot, and the slump structures and breccia of the plot belong to different earthquake rocks.

The linshuichonetes-Crurithyris community and new productid species from the Cisuralian (Early Permian) of Sichuan, China

Two marginiferinid species, Marginifera spinulifera sp. nov. and Transennatia sulcata sp. nov. are described from the Early Permian Liangshan and lowermost Chihsia formations at the Chuanmu Section, Sichuan, China. The Linshuichonetes – Crurithyris Community is discussed in the context of a pioneering opportunistic community that developed at the onset of the Yanghsingian transgression in the late Artinskian. This community was characterized by species that were generally small and relatively thin-shelled, and showed other features that are ecological adaptations to fluctuating environmental conditions.

Permian brachiopods from the Baoshan and Simao Blocks in Western Yunnan, China

This paper describes and illustrates four Permian brachiopod assemblages (Assemblages A–D) from western Yunnan, southwest China. Assemblage A occurs in the basal part of the Yongde Formation in the Xiaoxinzhai section in the southern Baoshan Block and is assignable to the latest Artinskian or earliest Kungurian. Assemblage B occurs about 50 m stratigraphically above Assemblage A in the same section, and is of Chihsian (Kungurian–Roadian) age. Assemblage C was collected from the Yongde Formation in the Anpaitian section in the southern Baoshan Block and is most likely Roadian to Wordian in age. Assemblage D came from the Longtan Formation in the Simao Block and is of late Wuchiapingian in age. The brachiopod faunas from the Yongde Formation of the Baoshan Block, as a whole, is dominated by species characteristic of the Cathaysian Province, although some links with Peri-Gondwanan faunas are also evident. In contrast, Assemblage D from the Simao Block is characterised exclusively by taxa of the Cathaysian Province. New taxa described and illustrated are Neochonetes (Huangichonetes) inflatus n. sp. Costatumulus minor n. sp. and Celebetus yunnanensis n. sp.

Permian brachiopods from the Tupe formation, San Juan Province, Precordillera, Argentina

The brachiopod fauna from the Tupe Formation at La Herradura Creek, located on the west flank of Perico Hill, San Juan Province, Argentina, palaeogeographically belongs to the western sector of the Paganzo basin ('Guandacol embayment'). The stratigraphical section of the Tupe Formation at La Herradura Creek is the stratotype of the Tivertonia jachalensis-Streptorhynchus inaequiornatus biozone, was previously regarded as being of Late Carboniferous age but here is assigned to the earliest Permian (Asselian). We describe and review the biozone assemblage, which consists of Streptorhynchus inaequiornatus, Tivertonia jachalensis, Kochiproductus sp., Costatumulus sp., Coronalosia argentinensis, Tupelosia paganzoensis, Trigonotreta pericoensis, Septosyringothyris sp. aff. Septosyringothyris jaguelensis and Crurithyris? sp. This brachiopod assemblage is related to Indian and Australian Early Permian faunas and its presence in the La Herradura Creek section provides new evidence in support of an Asselian (Early Permian) age for the Tivertonia jachalensis-Streptorhynchus inaequiornatus biozone. This assemblage is also important for intra- and inter-basinal correlation because several of its characteristic species have been identified from other sections of the Paganzo basin and the Riacuteo Blanco basin. The proposed age for this biozone is consistent with the age of palynological data from slightly above the marine faunas from the stratotype locality.

The Permian brachiopod genus Trigonotreta Koenig and its occurrence in Argentina

A review of the Permian spiriferid brachiopod genus Trigonolrela and its occurrence in the Early Permian of Argentina is provided herein. Several species are known from the Late Palaeozoic sequences of the Argentinean Precordillera. These include Trigonolrela sp. and Trigonolrela riojanensis (Lech and Acenolaza), from the Rio del Peii.on Formation (Rio Blanco Basin), Trigonolrela pericoensis (Leanza), from the Tupe Formation at the La Herradura creek locality (Paganzo Basin) and Trigonolrela sanjuanensis (Lech and Acenolaza), from Del Salto Formation (Calingasta-Uspallata Basin). These species are characterised by being small to medium sized, relatively transverse, with cardinal extremities often strongly angular. Costae are weakly bifurcated and superimposed on weakly developed lateral flank plications adjacent to the fastigium and sulcus. The Argentinean species are close to the oldest known Indian species of the genus, Trigonotreta hesdoensis (Salmi and Dutt), particularly with respect to the nature of its weakly fasciculated costae. Further study will refine the details of the relationship of the South American species with those from elsewhere in Gondwana and may permit the recognition of a distinctive lineage. The presence of the genus in Argentina in the earliest Permian is an important palaeobiogeographical observation that raises questions about the probable migration routes of the genus from the western Gondwanan South American margin to eastern Australia and India. The Precordilleran region appears to be the likely site of the first appearance of Trigonolrela. Species with relatively simple costae appeared first. These gave rise to more complex species with a more elaborate costal pattern indicating an evolutionary progression through time.

Late Paleozoic depositional history of the Tarim basin, northwest China: an integration of biostratigraphic and lithostratigraphic constraints

This study provides the first detailed lithostratigraphic and biostratigraphic constraints for improving stratigraphic resolution for hydrocarbon prospecting and exploration in the Tarim basin. A total of 49 stratigraphic units (38 formations and 11 members), ranging in age from the latest Devonian to Permian, are reviewed or redefined in terms of nomenclatures, lithology, age constraints, and lateral distributions based on the detailed field works or newly published data. Of these, the Piqiang Formation (new formation) is proposed to include the reefal carbonates of Asselian-Sakmarian age from the northern Tarim. The subsurface upper Paleozoic stratigraphic framework of the desert areas of the basin is also established for the first time. The high-resolution, basinwide stratigraphic correlations reveal that the sedimentation of the basin in the late Paleozoic was extremely uneven. Of these, the Famennian to Changhsingian successions are completely recorded in the south-western margin areas of the basin. Here, five eustatic sedimentary cycles are well recognizable, suggesting the sedimentation was more eustatically controlled and little affected by local tectonism. The late Paleozoic successions of both Kalpin and Taklimakan regions are commonly interrupted by major hiatuses at various horizons, suggesting that the sedimentation was apparently modified by local tectonism. Of these, the northward movement of the Tarim block and its subsequent collision with the Yili microcontinent (part of the Kazakhstan plate) may be principally accountable for the discrepancy in the sedimentation of the various regions in the basin in the late Paleozoic.

Unique marine Permian-Triassic boundary section from Western Australia

A unique marine Permian-Triassic boundary section containing rich oil source rocks has been continuously cored in a petroleum borehole from the Perth Basin of Western Australia. Such sequences, which provide a biostratigraphic and environmental record at the time of the largest extinction event of the past 500 million years, are globally rare, and this is the first to be documented in Australia. Throughout geological history there have been periods of global marine anoxia that commonly resulted in the widespread deposition of petroleum source rocks, most notably in the mid-Cretaceous and Late Jurassic. An apparent paradox is that, previously, source rocks have not been recognised in association with the Permian-Triassic boundary, despite widespread marine anoxia at this time. The Perth Basin source rocks contain abundant and unusual biomarkers, apparently related to the highly specialised and limited biota that flourished in the aftermath of the end-Permian extinction event. Local conditions may have favoured source-rock development, either due to higher productivity resulting from coastal upwelling or through enhanced preservation under strongly anoxic conditions.

Late palaeozoic global changes affecting high-latitude environments and biotas : an introduction

The Late Palaeozoic Ice Age (LPIA), spanning approximately from ~320 Ma (Serpukhovian, late Mississippian) to 290 Ma (mid-Sakmarian, Early Permian), represents the vegetated Earth’s largest and most long-lasting regime of severe and multiple glaciations, involving processes and patterns probably comparable to those of the Last Ice Age. Accompanying the LPIA occurred a number of broadly synchronous global environmental and biotic changes. These global changes, as briefly reviewed and summarized in this introductory paper, comprised (but are not limited to) the following: massive continental reorganization in the lead up to the final assembly of Pangea resulting in profound changes in global palaeogeography, palaeoceanography and palaeobiogeogarphy; substantially lowered global atmospheric carbon dioxide concentrations (pCO₂), coupled with an unprecedented increase in atmospheric oxygen concentrations reaching Earth's all-time high in its last 600 million year history; sharp global temperature and sea-level drops (albeit with considerable spatial and temporal variability throughout the ice age); and apparently a prolonged period of global sluggish macro-evolution with both low extinction and origination rates compared to other times. In the aftermath of the LPIA, the world's climate entered into a transitional climate phase through the late Early to Middle Permian before its transformation into a greenhouse state towards the end-Permian. In recent years, considerable amount of data and interpretations have been published concerning the physical evidence in support of the LPIA, its broad timeframe and eustatic and ecosystem responses from the lower latitudes, but relatively less attention has been drawn to the impact of the ice age on late Palaeozoic high-latitude environments and biotas. It is with this mission in mind that we have organized this special issue, with the central focus on late Palaeozoic high latitude regions of both hemispheres, that is, Gondwana and northern Eurasia. Our aim is to gather a set of papers that not only document the physical environmental changes that had occurred in the polar regions of Gondwana and northern Eurasia during the LPIA, but also review on the biotic responses at different taxonomic, ecological and spatial scales to these physical changes in a refined chronological timeframe.

This introductory paper is designed to provide a global context for the special issue, with a brief review of key late Palaeozoic global environmental changes (including: changes in global land-sea configurations, atmospheric chemistry, global climate regimes, global ocean circulation patterns and sea levels) and large -scale biotic (biogeographic and evolutionary) responses, followed by a summary of what we see as unresolved scientific issues and various working hypotheses concerning late Palaeozoic global changes and, in particular, the LPIA, as a possible reference to future research.

Implications of Kungurian (Early Permian) conodonts from Hatahoko, Japan, for correlation between the Tethyan and international timescales

The correlation between the fusulinid-based Tethyan and the conodont-based international timescales of the Permian System has become one of themost disputed issues among the Permian community during the past two decades. In this paper,we document a conodont fauna consisting of four species including Sweetognathus guizhouensis, Pseudohindeodus augustus, Hindeodus permicus and a new genus Meiognathus pustulus from the lower part of a large exotic limestone block at Hatahoko in the Nyukawa area, Gifu Prefecture, central Japan, which all suggest aKungurian age. The Kungurian age indicated by the conodonts is consistent with the age of the associated brachiopods, but conflicts with the Murgabian age indicated by the associated fusulinids including Cancellina nipponica, Neoschwagerina simplex, Neofusulinella praecursor etc. This co-occurrence of Kungurian conodonts and Murgabian fusulinids in central Japan suggests that previously unrecognized temporal distributions of some key fusulinid or conodont elements need to be clarified and that the intensively-disputed correlation problem between theKungurian containing theMurgabian fusulinids at the Luodian section in Guizhou, South China with the strata containing the ammonoid Waagenoceras in Oman and Sicily was caused by artificial conodont taxonomic discrepancies. The Luodian section in South China could serve as a key reference section for the correlation of the Kungurian Stage (late Early Permian) between the Tethyan and international timescales.