Information collected in Teesdale concerning free-swimming stages of brown trout (Salmo trutta L.)

During 1978 and 1979, electrofishing surveys were made in Teesdale - both to provide background information for ecological work on the streams, and to provide data so that the influence of discharge regime on the fish population densities could be examined. The discharge regimes of the different streams were compared using the Base Flow Index (BFI) as developed by the Institute of Hydrology. (PDF contains 30 pages)

Survival of intragravel stages of brown trout (Salmo trutta L.) in Teesdale

A study has been undertaken on several streams in Teesdale (UK)in order to examine survival rates of intragravel stages of brown trout and the factors influencing survival. Although all the becks contained brown trout spawning areas, some were utilised by more spawning trout than others. The best spawning sites as judged by this criterion were Thorsgill and Great Eggleshope becks where the research effort was therefore concentrated. There were two different spawning areas in Eggleshope, namely Great Eggleshope beck itself and Great Eggleshope spring fed tributary (Esft), data from these two areas are analysed separately in most parts of the text.

Mechanics of sediment transport and bedrock erosion in steep landscapes

Erosion is concentrated in steep landscapes such that, despite accounting for only a small fraction of Earth’s total surface area, these areas regulate the flux of sediment to downstream basins, and their rugged morphology records transient changes (or lack thereof) in geologic and climatic forcing. Steep landscapes are geomorphically active; large sediment fluxes and rapid landscape evolution rates can create or destroy habitat for humans and wildlife alike, and landslides, debris flows, and floods common in mountainous areas represent a persistent natural and structural hazard. Despite the central role that steep landscapes play in the geosciences and in landscape management, the processes controlling their evolution have been poorly studied compared to lower-gradient areas. This thesis focuses on the basic mechanics of sediment transport and bedrock incision in steep landscapes, as these are the fundamental processes which set the pace and style of landscape evolution. Chapter 1 examines the spatial distribution of slow-moving landslides; these landslides can dominate sediment fluxes to river networks, but the controls on their occurrence are poorly understood. Using a case-study along the San Andreas Fault, California, I show that slow-moving landslides preferentially occur near the fault, suggesting a rock-strength control on landslide distribution. Chapter 2 provides the first field-measurements of incipient sediment motion in streams steeper than 14% and shows a large influence of slope-dependent flow hydraulics and grain-scale roughness on particle motion. Chapter 3 presents experimental evidence for bedrock erosion by suspended sediment, suggesting that, in contrast to prevailing theoretical predictions, suspension-regime transport in steep streams can be the dominant erosion agent. Steep streams are often characterized by the presence of waterfalls and bedrock steps which can have locally high rates of erosion; Chapters 4 and 5 present newly developed, experimentally validated theory on sediment transport through and bedrock erosion in waterfall plunge pools. Finally, Chapter 6 explores the formation of a bedrock slot canyon where interactions between sediment transport and bedrock incision lead to the formation of upstream-propagating bedrock step-pools and waterfalls.

Ecological study of the River Lambourn. Report for the period October 1973 to September 1977

The investigations described in this report were carried out to fulfill three distinct but interrelated objectives. In 1973 the Thames Conservancy were making plans for a second stage of their groundwater scheme which would take water from the chalk aquifers in the valley of the Kennet and they wanted basic information on the ecological state of this river and its upper tributaries. Little appeared to be known about limestone streams and a preliminary study of one of the streams in this area was desirable as a basis for planning more detailed studies if these were needed later. At a progress meeting held in March 1976 the problems and opportunities presented by the developing drought conditions were considered. It was concluded that the ecological effects of the exceptionally low natural flows should be studied and that it would be important to assess the ecological impact of the groundwater scheme if it was brought into operation that year. This could only be done on the Lambourn and the Winterbourne and it was decided that considerable effort should be diverted there for this purpose and that the field observations should be extended to cover any recovery period after the end of the drought. To make this possible it was agreed that the studies of invertebrates and detritus on the Kennet should be reduced considerably and that the proposed study of the limestone stream should be abandoned. The revised objectives were as follows: A detailed ecological study of several sites on the Kennet and its tributaries above Kintbury, extending over at least two years and involving observations on wate r weeds , invertebrates, fish, detritus and the trophic relationships within the river community. Quantitativ e and qualitative sampling of water weeds and invertebrates during one year at a number of sites on several chalk streams to determine whether the patterns and relationships found in the Lambourn are also found at the other sites. Observations on the Lambourn at Bagnor were to continue for most of the period to look for long-term fluctuations and to enable these sites to act as controls with which the other sites could be compared. Further detailed studies on the Lambourn and the Winterbourne to assess the impact of low flows, trial pumping and the operation of the groundwater scheme.

Composition and bulk properties of Dorset river gravels

Considerable interest has been expressed in the composition of the stream gravels, the movement of bed materials and the relationship of sediment composition, packing and siltation of void space to invertebrate ecology. In the autumn of 1982, freeze-samples of gravel were obtained in Dorset streams. Data were required on the depth of salmonid egg pocke and were part of a broader investigation of regional variation in the independent variables of salmonid fish length, gravel size, current velocity and the resultant dependent variable ~egg burial depth. The Dorset river gravels examined are bimodal. The grain size distribution may be resolved into two near-normal frequency distributions interpreted as representing a primary framework or lattice of gravel particles into which a secondary matrix population of sand particles has penetrated.

Progress on the salmon investigation

An article detailing some of the conclusions of the salmon investigation undertaken by the author, on the River Eden and its tributaries, for the previous few years. It is proposed that seasonal changes in young salmon growth are related to water temperature variation. A figure is included showing length of fish compared to the average temperature of water in the River Eden over a two year duration. The article describes comparative work undertaken to date between three streams within the Thurso watershed and the River Eden. A table is included showing the average size of fish in each of the watercourses compared. Laboratory experiments on the effects of temperature on young salmon are outlined, as well as investigative work undertaken into the realtionship between fish scales and fish length.

Effects of injection nozzle configuration on secondary injection into supersonic flow

Gaseous nitrogen and argon were injected into a primary stream of air moving at Mach 2.56. The gases were injected at secondary to primary total pressure ratios from 3.2 to 28.6 through four different nozzles. Two nozzles, one sonic and one supersonic (M = 3.26), injected normal to the primary stream; and two sonic nozzles injected at 45° angles to the primary flow, one injecting upstream and the other downstream. Data consisted of static pressure measurements on the wall near the injector, total pressure profiles in the wake of the injectant plume, and concentration measurements downstream of the flow. Scale parameters were calculated based upon an analytical model of the flow field and their validity verified by experimental results. These scale heights were used to compare normalized wall side forces for the different nozzles and to compare the mixing of the two streams.

On the transport properties of fluid-particle flow

The hydrodynamic forces acting on a solid particle in a viscous, incompressible fluid medium at low Reynolds number flow is investigated mathematically as a prerequisite to the understanding of transport processes in two-phase flow involving solid particles and fluid. Viscous interaction between a small number of spherical particles and continuous solid boundaries as well as fluid interface are analyzed under a “point-force” approximation. Non-spherical and elastic spherical particles in a simple shear flow area are then considered. Non-steady motion of a spherical particle is briefly touched upon to illustrate the transient effect of particle motion.

A macroscopic continuum description of particle-fluid flow is formulated in terms of spatial averages yielding a set of particle continuum and bulk fluid equations. Phenomenological formulas describing the transport processes in a fluid medium are extended to cases where the volume concentration of solid particles is sufficiently high to exert an important influence. Hydrodynamic forces acting on a spherical solid particle in such a system, e.g. drag, torque, rotational coupling force, and viscous collision force between streams of different sized particles moving relative to each other are obtained. Phenomenological constants, such as the shear viscosity coefficient, and the diffusion coefficient of the bulk fluid, are found as a function of the material properties of the constituents of the two-phase system and the volume concentration of solid. For transient heat conduction phenomena, it is found that the introduction of a complex conductivity for the bulk fluid permits a simple mathematical description of this otherwise complicated process. The rate of heat transfer between particle continuum and bulk fluid is also investigated by means of an Oseen-type approximation to the energy equation.

Biological environments of larger UK rivers: progress report 1987

This interim progress report for the 9 months from January 1987 to September 1987 aims to provide insights into the mechanisms by which populations of particles (in this instance, phytoplankton) behave in relation to fluvial flow and, thus, to better model the dispersive properties of rivers and the ecological principles governing the distribution of potamoplankton generally. The author has been able to show with dye-tracers that significant water retention in pool reaches occurs within the range of (low) discharges obtaining, in accord with the Aggregated Dead Zone model and to an extent comparable with streams and small rivers investigated previously.

The production of algae growing on gravel in a chalk-stream

Epilithic algae, ie that growing on the surface of stones, was studied as part of the work on the energy flow of the chalk-stream ecosystem, by the River Laboratory. The study area was on Bere Stream and 2 neighbouring streams. The algal biomass was estimated from analysis of chlorophyll a. In Bere Stream the peak chlorophyll a cover occurred in April, while in the neighbouring streams, which have considerably lower nutrient levels, there was on peak. Assuming that 2% of a diatoms dry wt is chlorophyll a, then even in mid-April the biomass of epilithic algae amounted to no more than 15 g dry wt m Super(-2) of exposed gravel. Annual production was calculated to be > 15 times greater than biomass. The estimation of net primary production is always difficult for benthic floras and comparisons are especially difficult when different methods are used. But these figures contrast sharply with those for Ranunculus (water crowfoot) which has a ratio of annual production to maximal seasonal biomass of 1:16. The accumulation of algal biomass is apparently being prevented. Some organic matter may be excreted; some algae will be washed off the bed of the stream by current and grazing by herbivorous invertebrates will also tend to prevent algal accumulation.

The ecology of the Gussage, a lined winterbourne

The Gussage, a Dorset winterhourne (intermittent chalk stream), has been used to convey water from a compensation borehole to the River Allen to supplement its flow to meet demand for water. Sections of the Gussage have been lined with chalk, butyl sheeting or polythene sheeting to prevent water loss through the porous bed. The effects of this major environmental modification associated with these abstraction practices in the winterbourne catchments have been studied in the Gussage system since 1973. To compensate for the lack of adequate pre-lining data, comparative studies have been made on 3 small unlined chalk streams of varying flow regimes, ie. the Crichel (a winterbourne), the North Winterbourne and the Tarrant (permanent discharge in the reaches studies). The distribution of macrophytes and invertebrates in winterbournes are compared with that in natural and artificial permanent streams. Statistical analysis showed samples from the winterbourne sites and the unmodified permanent stream sites are quite distinct, despite the fact that no samples were taken from winterbourne sites during the dry phase. This emphasizes the differences between the fauna of an intermittent and a permanent stream and suggests that alteration of the flow regime could be a significant factor. Where flow regime has been altered, as in the Gussage downstream of the borehole, the samples occupy an intermediate position. Within this group of modified sites there is no apparent gross difference between the invertebrates of lined or unlined reaches.

Physiological aspects of the life cycle of the river lamprey, Lampetra fluviatilis L

The life cycle of the river lamprey, L. fluviatilis, is reviewed. The larval lamprey, or ammocoete, is a blind, filter-feeding animal, which normally lies concealed in the silt deposits of streams and rivers. After a period of 3-5 years in fresh water the ammocoete undergoes a metamorphosis in the summer months into a sexually immature, non-feeding stage known as the macrophthalia, which is active. This stage migrates downstream in late winter. It adopts a parasitic existence, in intertidal areas. After 18 months it returns to spawn in fresh water, after a final freshwater stage lasting up to 9 months. The river lamprey dies within a few days after the spawning period of 3-4 weeks, and none survive to spawn the following year.

Acid precipitation and its effects on aquatic systems in the English Lake District

There is no evidence of an increase in the acidity (lower pH or alkalinity) of water-bodies in the Lake District over the last 50 years. Brown trout occur in acid streams and upland tarns where pH is 4.5-5.2 throughout the year. Their occurrence in such waters in Britain and Ireland has been known for most of this century and there is no previous evidence of harmful effects on salmonid fisheries, though numbers of fish are naturally low. However, many benthic invertebrates that are common in hill-streams where pH is above 5.7 do not occur in more acid streams. This phenomenon occurs in the headwaters of several western rivers in Cumbria. It is not a recent response to "acid rain". Harmful effects of pH are undoubtedly more pronounced in waters that are poor in other dissolved ions. Low concentrations of sodium, potassium, calcium and chloride are especially important and may limit the distributions of some aquatic animals even where pH is above 5.7. The concentration of sulphate ions is usually relatively high but this is not important to the fauna; concentrations are at least two times higher in productive alkaline water-bodies than they are in unproductive acid waters.

Experiments using an artificial stream to investigate the seasonal growth of chalk-stream algae

Research into the production ecology of chalk streams using a large artificial recirculating stream is described. Physical chemical processes including calcium and inorganic phosphate levels, and exchange of gaseous carbon dioxide in both a simple closed system and a circulating system with gravel substrate have been monitored in both light and dark conditions. Further experiments were concerned with the seasonal changes in algal growth over the gravel substrate with constant water velocities and replenishment. The algal population, composed mainly of the diatoms Achnanthes minutissima, Meridion circulare, Nitzschia fonticola and Synedra ulna reached a peak in mid May and declined rapidly during June. Concentrations of phosphate phosphorus fell as the diatoms grew but was not thought to limit growth. Silicate concentrations followed the diatom cycle closely but never fell below 0.8 mg/l Si. It is possible that one of the nutrients may have been limiting the rate of growth due to steep diffusion gradients through the algal mat. In the last summer and autumn a hard calcareous crust composed of the green alga Gongrosira incrustans and the blue green alga Homeothrix varians , developed. The channel stream is compared with the natural conditions found in chalk streams.

The ecology of chalk-stream invertebrates studied in a recirculating stream

To study and qualify the factors influencing interactions between various trophic levels in natural hard-water streams, a recirculating artificial stream channel was constructed. This structure has enabled patterns of population change of stream fauna to be observed under partially controlled physical and chemical conditions. Initial colonization of the substratum by invertebrates and subsequent succession was studied along with depth distribution and growth and production studies of invertebrates.