Phylogenetic relationships of the hoatzin, an enigmatic South American bird.

The hoatzin (Opisthocomus hoazin) lives in the humid lowlands of northern and central South America, often in riparian habitats. It is a slender bird approximately 65 cm in length, brownish with lighter streaks and buffy tips to the long tail feathers. The small head has a ragged, bristly crest of reddish-brown feathers, and the bare skin of the face is bright blue. It resembles a chachalaca (Ortalis, Cracidae) in size and shape, but its plumage and markings are similar to those of the smaller guira cuckoo (Guira guira). The hoatzin (pronounced Watson) has been a taxonomic puzzle since it was described in 1776. It usually has been viewed as related to the gallinaceous birds, but alliances to other groups have been suggested, including the cuckoos. We present DNA sequence evidence from the 12S and 16S rRNA mitochondrial genes, and from the nuclear gene that codes for the eye lens protein, alpha A-crystallin. The results indicate that the hoatzin is most closely related to the typical cuckoos and that the divergence occurred at or near the base of the cuculiform phylogenetic tree.

Central & Eastern Africa, 1635 (Raster Image)

This layer is a georeferenced raster image of the historic paper map entitled: Aethiopia superior vel interior, vulgo Abissinorum sive Presbiteri Ioannis imperium. It was published by Guiljelmum et Johannem Blaeu in 1635. Scale [ca. 1:12,600,000]. Covers Central and Eastern Africa. Map in Latin.The image inside the map neatline is georeferenced to the surface of the earth and fit to the Africa Sinusoidal projected coordinate system. All map collar and inset information is also available as part of the raster image, including any inset maps, profiles, statistical tables, directories, text, illustrations, index maps, legends, or other information associated with the principal map. This map shows features such as drainage, cities and other human settlements, territorial boundaries, shoreline features, and more. Relief shown pictorially. Includes notes, illustrations of animals, and ornamental cartouche. This layer is part of a selection of digitally scanned and georeferenced historic maps from the Harvard Map Collection. These maps typically portray both natural and manmade features. The selection represents a range of originators, ground condition dates, scales, and map purposes.

Central & Eastern Africa, 1655 (Raster Image)

This layer is a georeferenced raster image of the historic paper map entitled: Haute Ethiopie, ou sont L'Empire des Abissins, La Nubie, et le Zanguebar : subdivisés en leurs principales parties, tirés de Sanut de Mercator &c. par le Sr. Sanson d'Abbeville. It was published by P. Mariette in 1655. Covers Central and Eastern Africa. Scale [ca. 1:12,000,000]. Map in French.The image inside the map neatline is georeferenced to the surface of the earth and fit to the Africa Sinusoidal projected coordinate system. All map collar and inset information is also available as part of the raster image, including any inset maps, profiles, statistical tables, directories, text, illustrations, index maps, legends, or other information associated with the principal map. This map shows features such as drainage, cities and other human settlements, territorial boundaries, shoreline features, and more. Relief shown pictorially.This layer is part of a selection of digitally scanned and georeferenced historic maps from the Harvard Map Collection. These maps typically portray both natural and manmade features. The selection represents a range of originators, ground condition dates, scales, and map purposes.

Central & Eastern Africa, 1887 (Raster Image)

This layer is a georeferenced raster image of the historic paper map entitled: Central Africa : on a scale of 1:10,000,000, By Dr. F. Boas. It was published by Hart & Von Arx in 1887. Scale 1:10,000,000 The image inside the map neatline is georeferenced to the surface of the earth and fit to the Africa Sinusoidal projected coordinate system. All map collar and inset information is also available as part of the raster image, including any inset maps, profiles, statistical tables, directories, text, illustrations, index maps, legends, or other information associated with the principal map. This map shows features such as drainage, cities and other human settlements, shoreline features, and more. Relief shown by shading. This layer is part of a selection of digitally scanned and georeferenced historic maps from the Harvard Map Collection. These maps typically portray both natural and manmade features. The selection represents a range of originators, ground condition dates, scales, and map purposes.

Cape Town, South Africa, 1911 (Raster Image)

This layer is a georeferenced raster image of the historic paper map entitled: Map of Cape Town (Central). It was published by Cape Peninsula Publicity Association in 1911. Scale [ca. 1:4,700]. Covers a portion of Cape Town, South Africa. The image inside the map neatline is georeferenced to the surface of the earth and fit to the Universal Transverse Mercator (UTM Zone 34S, meters, WGS 1984) projected coordinate system. All map collar and inset information is also available as part of the raster image, including any inset maps, profiles, statistical tables, directories, text, illustrations, index maps, legends, or other information associated with the principal map. This map shows features such as roads, railroads and stations, street-railroads, drainage, selected buildings and tourist attractions, parks, and more. Includes inset location map and an index to points of interest.This layer is part of a selection of digitally scanned and georeferenced historic maps from the Harvard Map Collection. These maps typically portray both natural and manmade features. The selection represents a range of originators, ground condition dates, scales, and map purposes.

China, North Korea, and South Korea, 1749 (Raster Image)

This layer is a georeferenced raster image of the historic paper map entitled: La Chine avec la Korée, et les parties de la Tartarie les plus voisines : tirées des cartes que les Jesuites Missionaires ont levées les années 1708, jusqu'en 1717, J. V. Schley direx. It was published by Pierre de Hondt in 1749. Scale [ca. 1:2,100,000]. Covers East and Central China, and North and South Korea, and a small portion of Mongolia. Map in French and Dutch.The image inside the map neatline is georeferenced to the surface of the earth and fit to the Asia North Lambert Conformal Conic coordinate system. All map collar and inset information is also available as part of the raster image, including any inset maps, profiles, statistical tables, directories, text, illustrations, index maps, legends, or other information associated with the principal map. This map shows features such as drainage, cities and other human settlements, territorial boundaries, shoreline features, the Great Wall of China, and more. Relief shown pictorially. This layer is part of a selection of digitally scanned and georeferenced historic maps from the Harvard Map Collection. These maps typically portray both natural and manmade features. The selection represents a range of originators, ground condition dates, scales, and map purposes.

China, North Korea, and South Korea, 1808 (Raster Image)

This layer is a georeferenced raster image of the historic paper map entitled: China, Neele sculpt. It was published by Longman & Co, Paternoster Row in Jany. 1, 1808. Scale [ca. 1:9,600,000]. Covers East and Central China, and North and South Korea, and a small portion of Mongolia, Vietnam, Laos, Burma, India, Japan, and Russia. Map in French and Dutch. The image inside the map neatline is georeferenced to the surface of the earth and fit to the Asia North Lambert Conformal Conic coordinate system. All map collar and inset information is also available as part of the raster image, including any inset maps, profiles, statistical tables, directories, text, illustrations, index maps, legends, or other information associated with the principal map. This map shows features such as drainage, cities and other human settlements, territorial boundaries, shoreline features, and more.This layer is part of a selection of digitally scanned and georeferenced historic maps from the Harvard Map Collection. These maps typically portray both natural and manmade features. The selection represents a range of originators, ground condition dates, scales, and map purposes.

East and Central China, 1841 (Raster Image)

This layer is a georeferenced raster image of the historic paper map entitled: Karte von China mit dessen Eintheilung in 18 Provinzen, nach den neuesten u. zuverlässigsten Materialien entworfen, gezeichnet und bearbeitet von J. B. Roost ; in Stein grawirt v. D. Grasmüller. It was published der liter. artist. Anstalt der L.G. Cotta'schen Buchhandlung in 1841. Scale 1:6,500,000. Covers East and Central China, and the Korean Peninsula. Map in German. The image inside the map neatline is georeferenced to the surface of the earth and fit to the Asia North Lambert Conformal Conic coordinate system. All map collar and inset information is also available as part of the raster image, including any inset maps, profiles, statistical tables, directories, text, illustrations, index maps, legends, or other information associated with the principal map. This map shows features such as drainage, cities and other human settlements, roads, territorial and provincial boundaries, shoreline features, and more.This layer is part of a selection of digitally scanned and georeferenced historic maps from the Harvard Map Collection. These maps typically portray both natural and manmade features. The selection represents a range of originators, ground condition dates, scales, and map purposes.

South America, 1838 (Raster Image)

This layer is a georeferenced raster image of the historic paper map entitled: Carte de l'Amérique Méridionale, dressée par Alcide d'Orbigny d'aprés ses itinéraires sur parties qu'il à parcourues servant à l'intelligence de son voyage et de ses observations sur l'homme américain ; gravé sur pierre par L. Bouffard ; lith de Lemercier Benard et Cie. It was published by Pitois-Levrault et Cie Libraire in 1838. Scale [ca. 1:15,000,000]. Covers South America. Map in French. The image inside the map neatline is georeferenced to the surface of the earth and fit to a non-standard 'World Sinusoidal' projection with the central meridian at 57.662770 degrees west. All map collar and inset information is also available as part of the raster image, including any inset maps, profiles, statistical tables, directories, text, illustrations, index maps, legends, or other information associated with the principal map. This map shows features such as drainage, roads, cities and other human settlements, cultural territories, selected land and sea exploration routes, shoreline features, and more. Relief shown by hachures. Includes inset: Bolivia d'après la carte Lapie (1829). This layer is part of a selection of digitally scanned and georeferenced historic maps from the Harvard Map Collection and the Harvard University Library as part of the Open Collections Program at Harvard University project: Organizing Our World: Sponsored Exploration and Scientific Discovery in the Modern Age. Maps selected for the project correspond to various expeditions and represent a range of regions, originators, ground condition dates, scales, and purposes.

South America, 1902 (Raster Image)

This layer is a georeferenced raster image of the historic paper map entitled: Rand, McNally & Co.'s new 14 x 21 map of South America. It was published by Rand, McNally & Co. in 1902. Scale [1:19,008,000]. The image inside the map neatline is georeferenced to the surface of the earth and fit to a non-standard 'World Sinusoidal' projection with the central meridian at 60 degrees west. All map collar and inset information is also available as part of the raster image, including any inset maps, profiles, statistical tables, directories, text, illustrations, index maps, legends, or other information associated with the principal map. This map shows features such as drainage, cities and other human settlements, territorial boundaries, shoreline features, and more. Relief shown by hachures. Includes inset of the Galapagos Islands. This layer is part of a selection of digitally scanned and georeferenced historic maps from the Harvard Map Collection and the Harvard University Library as part of the Open Collections Program at Harvard University project: Organizing Our World: Sponsored Exploration and Scientific Discovery in the Modern Age. Maps selected for the project correspond to various expeditions and represent a range of regions, originators, ground condition dates, scales, and purposes.

South America, 1739 (Image 1 of 2) (Raster Image)

This layer is a georeferenced raster image of the historic paper map entitled: Carte du Perou, pour servir à l'histoire des Incas et à celle de l'etat present de cette province, dressée par Philippe Buache, sur les observations astronomiq[es] faites aux environs de l'équateur et communiquées à l'Académie depuis l'an 1736 jusqu'en 1739 par M.M. Godin, Bouguer et de la Condaminede l'Acad. R[le] des Sciences Assujetie por les autres parties aux observations du P. Feuillée et de M. Frezier aux routes et remarques geographiques de divers voyageurs. It is part of a two sheet map set: [Carte du Peru]. It was published sur le Quay de la Megisserie avec privilege du Roy in 1739. Scale [ca. 1:195,000]. Covers the territory of the former Inca Empire, South America. This layer is image 1 of 2 total images of the two sheet source map representing the northern portion of the map. Map in French and Spanish. The image inside the map neatline is georeferenced to the surface of the earth and fit to a non-standard 'World Sinusoidal' projection with the central meridian at 75 degrees west. All map collar and inset information is also available as part of the raster image, including any inset maps, profiles, statistical tables, directories, text, illustrations, index maps, legends, or other information associated with the principal map. This map shows features such as drainage, cities and other human settlements, territorial boundaries, shoreline features, and more. Relief shown pictorially. This layer is part of a selection of digitally scanned and georeferenced historic maps from the Harvard Map Collection and the Harvard University Library as part of the Open Collections Program at Harvard University project: Organizing Our World: Sponsored Exploration and Scientific Discovery in the Modern Age. Maps selected for the project correspond to various expeditions and represent a range of regions, originators, ground condition dates, scales, and purposes.

South America, 1739 (Image 2 of 2) (Raster Image)

This layer is a georeferenced raster image of the historic paper map entitled: Carte de la partie meridionale du Perou : pour servir à l'histoire des Incas et à celle de l'etat present de cette province, dressée par Philippe Buache. It is part of a two sheet map set: [Carte du Peru]. It was published sur le Quay de la Megisserie avec privilege du Roy in 1739. Scale [ca. 1:195,000]. Covers the territory of the former Inca Empire, South America. This layer is image 2 of 2 total images of the two sheet source map representing the southern portion of the map. Map in French and Spanish. The image inside the map neatline is georeferenced to the surface of the earth and fit to a non-standard 'World Sinusoidal' projection with the central meridian at 75 degrees west. All map collar and inset information is also available as part of the raster image, including any inset maps, profiles, statistical tables, directories, text, illustrations, index maps, legends, or other information associated with the principal map. This map shows features such as drainage, cities and other human settlements, territorial boundaries, shoreline features, and more. Relief shown pictorially. This layer is part of a selection of digitally scanned and georeferenced historic maps from the Harvard Map Collection and the Harvard University Library as part of the Open Collections Program at Harvard University project: Organizing Our World: Sponsored Exploration and Scientific Discovery in the Modern Age. Maps selected for the project correspond to various expeditions and represent a range of regions, originators, ground condition dates, scales, and purposes.

South America, 1779 (Raster Image)

This layer is a georeferenced raster image of the historic paper map entitled: Carte du Pérou et Brésil sepl. de Tierre - Firme, de Guayana, et de la rivière des Amazones : ce qui fait la partie boréale de L'Amérique Meridiole., par le sr. d'Anville. It was published by P. Santini, Chez M. Remondini in 1779. Scale [ca. 1:6,500,000]. Covers the northern part of South America. Map in French. The image inside the map neatline is georeferenced to the surface of the earth and fit to a non-standard 'World Sinusoidal' projection with the central meridian at 58.000000 degrees west. All map collar and inset information is also available as part of the raster image, including any inset maps, profiles, statistical tables, directories, text, illustrations, index maps, legends, or other information associated with the principal map. This map shows features such as drainage, cities and other human settlements, territorial boundaries, shoreline features, and more. Relief is shown pictorially. This layer is part of a selection of digitally scanned and georeferenced historic maps from the Harvard Map Collection and the Harvard University Library as part of the Open Collections Program at Harvard University project: Organizing Our World: Sponsored Exploration and Scientific Discovery in the Modern Age. Maps selected for the project correspond to various expeditions and represent a range of regions, originators, ground condition dates, scales, and purposes.