Pollen profile of sediment core Dethlingen, Germany

To better understand the environmental variability during the Holsteinian interglacial, we have palynologically analyzed a new core from Dethlingen, northern Germany, at a decadal resolution. Our data provide insights into the vegetation dynamics and thus also climate variability during the meso- to telocratic forest phases of the interglacial. Temperate mixed forests dominated the regional landscape throughout the Holsteinian. However, changes in the forest composition during the younger stages of the interglacial suggest a climatic transition towards milder conditions in winter. The strong presence of boreal floral elements during the older stages of the Holsteinian interglacial suggests a high seasonality. In contrast, during the younger stages the development of sub-Atlantic and Atlantic floral elements suggests increasingly warm and humid climatic conditions. Peak warming during the younger stage of the Holsteinian is marked by the maximum pollen abundances of Buxus, Abies, and Quercus. Although the vegetation dynamics suggest a general warming trend throughout the Holsteinian interglacial, abrupt as well as gradual changes in the relative abundances of temperate plants indicate considerable climatic variability. In particular, two marked declines in temperate taxa leading to the transient development of boreal and sub-temperate forests indicate short-term climatic oscillations that occurred within full interglacial conditions. The palynological signatures of these two regressive phases in vegetation development differ with regard to the expansion of pioneer trees, the abundances and rates of change of temperate taxa, and the presence of frost-sensitive taxa. These differences point to different mechanisms responsible for the individual regressive phases. Assuming a correlation of the interglacial at Dethlingen with Marine Isotope Stage (MIS) 11, our data suggest that temperate forests prevailed in northern Germany during the younger parts of MIS 11c.

(Fig. 9) Pollen analysis of sediment core GIK16017-2

The improved understanding of the pollen signal in the marine sediments offshore of northwest Africa is applied to deep-sea core M 16017-2 at 21°N. Downcore fluctuations in the percentage, concentration and influx diagrams record latitudinal shifts of the main northwest African vegetation zones and characteristics of the trade winds and the African Easterly Jet. Time control is provided by 14C ages and 180 records. During the period 19,000-14,000 yr B.P. a compressed savanna belt extended between about 12 ° and 14-15°N. The Sahara had maximally expanded northward and southward under hyperarid climatic conditions. The belt with trade winds and dominant African Easterly Jet transport had not shifted latitudinally. The trade winds were strong as compared to the modern situation but around 13,000 yr B.P. the trade winds weakened. After 14,000 yr B.P. the climate became less arid south of the Sahara and a first spike of fluvial runoff is registered around 13,000 yr B.P. Fluvial runoff increased strongly around 11,000 yr B.P. and maximum runoff is recorded from about 9000-7800 yr B.P. Around 12,500 yr B.P. the savanna belt started to shift northward and became richer in woody species: it shifted about 6° of latitude, reached its northernmost position during the period of 9200-7800 yr B.P. and extended between about 16° and 24°N at that time. Tropical forest had reached its maximum expansion and the Guinea zone reached as far north as about 15°N, reflecting very humid climatic conditions south of the Sahara. North of the Sahara the climate also became more humid and Mediterranean vegetation developed rapidly. The Sahara had maximally contracted and the trade winds were weak and comparable with the present day intensity. After about 7800 yr B.P. the southern fringe of the Sahara and accordingly the savanna belt, shifted rapidly southward again.

Palynological and stratigraphic record from Löddigsee in Mecklenburg-Vorpommern, Germany

A Late Pleistocene and Holocene sediment core from the nowadays terrestrialised portion of the Löddigsee in Southern Mecklenburg, Germany was palynologically investigated. The lake is situated in the rarely investigated Young moraine area at the transition from the Weichselian to the Saalian glaciation. The high-resolution pollen diagram contributes to the establishment of the north-eastern German Late Pleistocene pollen stratigraphy. The vegetation distribution pattern after the end of the Weichselian is in good agreement with other studies from North-eastern Germany, but also has its own characteristics. The Holocene vegetation development reveals features from the north-eastern and north-western German lowlands. A special focus was laid on the environmental history of the two settlements on an island within the lake (Late Neolithic and Younger Slavic period), which were preserved under moist conditions. Both settlements were constructed during a period of low lake level. Although there is evidence of agriculture in the area during the respective periods, the two island settlements seem to have served other purposes.

Physoderma pulposum; Physodermataceae

Physoderma pulposum Wallr.

Zoocoenological state of microhabitats and its seasonal dynamics in an aquatic macroinvertebrate assembly (Hydrobiological case studies on lake Balaton, No. 1.)

In the years 2002, 2003 and 2004 we collected samples of macroinvertebrates on a total of 36 occasions in Badacsony bay, in areas of open water (in the years 2003 and 2004 reed-grassy) as well as populated by reed (Phragmites australis) and cattail (Typha angustifolia). Samples were taken using a stiff hand net. The sampling site includes three microhabitats differentiated only by the aquatic plants inhabiting these areas. Our data was gathered from processing 208 individual samples. The quantity of macroinvertebrates is represented by biovolume value based on volume estimates. We can identify taxa in abundant numbers found in all water types and ooze; as well as groups associated with individual microhabitats with various aquatic plants. We can observe a notable difference between the years in the volume of invertebrate macrofauna caused by the drop of water level, and the multiplication of submerged macrophytes. There are smaller differences between the samples taken in reeds and cattail stands. In the second half of 2003 – which was a year of drought – the Najas marina appeared in open waters and allowed to support larger quantities of macroinvertebrates. In 2004 with higher water levels, the Potamogeton perfoliatus occurring in the same area has had an even more significant effect. This type of reed-grass may support the most macroinvertebrates during the summer. From the aspect of diversity relations we may suspect different characteristics. The reeds sampling site proved to be the richest, while the cattail microhabitat is close behind, open water (with submerged macrophytes) is the least diverse microhabitat.

Seasonal dynamics of an aquatic macroinvertebrate assembly (Hydrobiological case study of Lake Balaton No. 2)

In 2002, 2003 and 2004, we took macoinvertebrate samples on a total of 36 occasions at the Badacsony bay of Lake Balaton. Our sampling site was characterised by areas of open water (in 2003 and 2004 full of reed-grass) as well as by areas covered by common reed (Phragmites australis) and narrowleaf cattail (Typha angustifolia). Samples were taken both from water body and benthic ooze by use of a stiff hand net. We have gained our data from processing 208 individual samples. We took samples frequently from early spring until late autumn for a deeper understanding of the processes of seasonal dynamics. The main seasonal patterns and temporal changes of diversity were described. We constructed a weather-dependent simulation model of the processes of seasonal dynamics in the interest of a possible further utilization of our data in climate change research. We described the total number of individuals, biovolume and diversity of all macroinvertebrate species with a single index and used the temporal trends of this index for simulation modelling. Our discrete deterministic model includes only the impact of temperature, other interactions might only appear concealed. Running the model for different climate change scenarios it became possible to estimate conditions for the 2070-2100 period. The results, however, should be treated very prudently not only because our model is very simple but also because the scenarios are the results of different models.

Enzyme Differences between Adjacent Hybrid and Parent Populations of Typha

Enzyme variation among three adjacent Typha populations was analyzed by means of disc electrophoresis, isoelectric focusing, and enzyme assays. Esterase isozyme analysis of seedlings of Typha latifolia L.,T . x glauca Godr., and T. angustifolia L. indicated that there was little or no variation within each population. Additional analyses of esterase, malate dehydrogenase, glutamate dehydrogenase, and alcohol dehydrogenase of pooled samples of pollen and seedlings indicated that the Typha x glauca population was the product of hybridization between the adjacent parent populations. The hybrid population had more isozymes, but lower specific activities, than the parents.

A Serological and Disc Electrophoretic Study of North American Typha

Pollen and seed proteins of seven selected North American and Puerto Rican Typha populations were compared using two serological methods and disc electrophoresis. These methods were capable of discriminating among all taxa studied: Typha latifolia, T. angustifolia, T. X glauca, and T. domingensis. The two hybrid populations were found to contain proteins not found in either parent. Typha domingensis was serologically the most distinct of the four taxa. The diagnostic morphological characteristics for Typha species were studied in all populations, and statistical comparisons are presented. Data from the morphological observations agreed with the information obtained from the chemosystematic research. All data indicate that the three taxa should be maintained as separate species. The hybrid nature of the putative T. X glauca is verified by both the biochemical and morphological data. Observed morphological and biochemical differences support taxonomic treatments in which T. domingensis is designated as a separate species.

Pollen and dinoflagellate cyst counts on sediment core M72/5_628-1 (MD72/5-25-GC1) from the Black Sea

High-resolution pollen and dinoflagellate cyst records from sediment core M72/5-25-GC1 were used to reconstruct vegetation dynamics in northern Anatolia and surface conditions of the Black Sea between 64 and 20 ka BP. During this period, the dominance of Artemisia in the pollen record indicates a steppe landscape and arid climate conditions. However, the concomitant presence of temperate arboreal pollen suggests the existence of glacial refugia in northern Anatolia. Long-term glacial vegetation dynamics reveal two major arid phases ~64-55 and 40-32 ka BP, and two major humid phases ~54-45 and 28-20 ka BP, correlating with higher and lower summer insolation, respectively. Dansgaard-Oeschger (D-O) cycles are clearly indicated by the 25-GC1 pollen record. Greenland interstadials are characterized by a marked increase in temperate tree pollen, indicating a spread of forests due to warm/wet conditions in northern Anatolia, whereas Greenland stadials reveal cold and arid conditions as indicated by spread of xerophytic biomes. There is evidence for a phase lag of ~500 to 1500 yr between initial warming and forest expansion, possibly due to successive changes in atmospheric circulation in the North Atlantic sector. The dominance of Pyxidinopsis psilata and Spiniferites cruciformis in the dinocyst record indicates brackish Black Sea conditions during the entire glacial period. The decrease of marine indicators (marine dinocysts, acritarchs) at ~54 ka BP and increase of freshwater algae (Pediastrum, Botryococcus) from 32 to 25 ka BP reveals freshening of the Black Sea surface water. This freshening is possibly related to humid phases in the region, to connection between Caspian Sea and Black Sea, to seasonal freshening by floating ice, and/or to closer position of river mouths due to low sea level. In the southern Black Sea, Greenland interstadials are clearly indicated by high dinocyst concentrations and calcium carbonate content, as a result of an increase in primary productivity. Heinrich events show a similar impact on the environment in the northern Anatolia/Black Sea region as Greenland stadials.

Pollen distribution of sediment core MD96-2048 from the Western Indian Ocean

Glacial-interglacial fluctuations in the vegetation of South Africa might elucidate the climate system at the edge of the tropics between the Indian and Atlantic Oceans. However, vegetation records covering a full glacial cycle have only been published from the eastern South Atlantic. We present a pollen record of the marine core MD96-2048 retrieved by the Marion Dufresne from the Indian Ocean ~120 km south of the Limpopo River mouth. The sedimentation at the site is slow and continuous. The upper 6 m (spanning the past 342 Ka) have been analysed for pollen and spores at millennial resolution. The terrestrial pollen assemblages indicate that during interglacials, the vegetation of eastern South Africa and southern Mozambique largely consisted of evergreen and deciduous forests. During glacials open mountainous scrubland dominated. Montane forest with Podocarpus extended during humid periods was favoured by strong local insolation. Correlation with the sea surface temperature record of the same core indicates that the extension of mountainous scrubland primarily depends on sea surface temperatures of the Agulhas Current. Our record corroborates terrestrial evidence of the extension of open mountainous scrubland (including fynbos-like species of the high-altitude Grassland biome) for the last glacial as well as for other glacial periods of the past 300 Ka.

Pollen profile and age determination for sediment core M72/5_619-1 (22-GC3)

Sediments from the Black Sea, a region historically dominated by forests and steppe landscapes, are a valuable source of detailed information on the changes in regional terrestrial and aquatic environments at decadal to millennial scales. Here we present multi-proxy environmental records (pollen, dinoflagellate cysts, Ca, Ti and oxygen isotope data) from the uppermost 305 cm of the core 22-GC3 (42°13.53' N, 36°29.55' E) collected from a water depth of 838 m in the southern part of the Black Sea in 2007. The records span the last ~ 18 kyr (all ages are given in cal kyr BP). The pollen data reveal the dominance of the Artemisia-steppe in the region, suggesting rather dry/cold environments ~ 18-14.5 kyr BP. Warming/humidity increase during melt-water pulses (~ 16.1-14.5 kyr BP), indicated by d18O records from the 22-GC3 core sediment and from the Sofular Cave stalagmite, is expressed in more negative d13C values from the Sofular Cave, usually interpreted as the spreading of C3 plants. The records representing the interstadial complex (~ 14.5-12.9 kyr BP) show an increase in temperature and moisture, indicated by forest development, increased primary productivity and reduced surface run-off, whereas the switch from primary terrigenous to primary authigenic Ca origin occurs ~ 500 yr later. The Younger Dryas cooling is clearly demonstrated by more negative d13C values from the Sofular Cave and a reduction of pines. The early Holocene (11.7-8.5 kyr BP) interval reveals relatively dry conditions compared to the mostly moist and warm middle Holocene (8.5-5 kyr BP), which is characterized by the establishment of the species-rich warm mixed and temperate deciduous forests in the low elevation belt, temperate deciduous beech-hornbeam forests in the middle and cool conifer forest in upper mountain belt. The border between the early and middle Holocene in the vegetation records coincides with the opening of the Mediterranean corridor at ~ 8.3 kyr BP, as indicated by a marked change in the dinocyst assemblages and in the sediment lithology. Changes in the pollen assemblages indicate a reduction in forest cover after ~ 5 kyr BP, which was likely caused by increased anthropogenic pressure on the regional vegetation.