Benthos of Spanish lakes and reservoirs

A summary of the results from the study of benthos of lakes and reservoirs in Spain is provided, with a list of the species found to date. Spanish natural lakes are smaller than European lakes; the largest is Lake Sanabria, of glacial origin, which is 3 Km long and half a kilorneter wide. Many are very small and situated in the mountains; more than 200 hundred have been recorded in Spain, but only in Lake Sanabria and Lake Banyoles have the benthos been studied. Lake Sanabria is a cold oligotrophic, monomictic lake with oxygen always present in the deepest zones. Its fauna is similar to that of other central European lakes, with Mici.opsectra c.orztractu (a chironomid) as the dominan1 species. Lake Banyoles is a karstic, monomictic and multibasin lake; despite the low primary productivity, due to the abundante of sulphate in the water, the allochthonous inflow of organic matter and the inflow of water from bottom springs, the profunda1 environinent is very stressing for benthic fauna. Very low oxygen concentrations and high sulphide content in the water and sediments dueto meromixis mean that only the larva of the dipteran Chaohoi.lcs flai7icans was present in one of the 5 basins of the lake. In other basins, when oxygen is available (no meromixis), the fauna is similar to that of tlie inineralized lakes of the Aegean region and some lakes in central Italia. On the other hand, preliminary data from the Pyrenean lakes and from Sierra Nevada ponds reveal no differences with northern cold lakes. Tlie largest lakes in Spain are the reservoirs. There are nearly 1000 and data are available on 100 of them, including the kargest. In addition to oxygen and sulphide content in the bottom waters, water level fluctuation and high sedimentation rates are disturbance factors that prevented the organization of the community. Allochthonous inputs of organic matter are also an important factor both in the reservoirs and also in the small, oligotrophic lakes like Banyoles and Sanabria. As a result the meiofaunal loop is very important in many of the Spanish water bodies . For this reason the natural lakes and reservoirs of Spain are dominated by Oligochaeta, small crustaceans and the microcarnivore chironomids (such as Procladi~ls, Cladopelma and Mi(,rnc.hil-onnmus) that feed on these meoifaunal elements. The phytophagous chironomids, like Chironomus, are only abundan1 in the shallow areas of mesotrophic and eutrophic reservoirs. This situation makes it difficult to apply the typological system of SAETHER which predicts with some confidence only the benthic communities of Spanish natural lakes above 1500 m in the Pyrenees or the ponds above 2000 m in Sierra Nevada mountains. Higher temperatures (which originate a longer stratification period), the presence of sulphate in the waters of the eastern part of Spain and high inputs of sediments and allochthonous organic matter seem to be the factors that originated the differences between the benthic profunda1 faunas of Spanish lakes and reservoirs and those of the temperate lakes of north and central Europe.

Pulmonaria longifolia (Bast.) Boreau subsp. longifolia al massis de Garraf

Pulmonaria

Observaciones sobre la colonizacion briologico-liquenica de la madera en descomposicion en los bosques del Moixero (Catalunya)

Con este estudio se pretende iniciar una serie de trabajos pera conocer la flora briológico-liquénica que coloniza la madera en descomposición. Exponemos en este artículo los resultados obtenidos en los bosques septentrionales del Moixeró. Así mismo indicamos la posible sucesión existente en la colonización del substrato según su grado de descomposición. Esta sucesión se ha descrito a partir de las observaciones realizadas en tocones y troncos en distintos grados de descomposición.

Phylogeny and biogeography of Primula sect. Armerina: implications for plant evolution under climate change and the uplift of the Qinghai-Tibet Plateau.

BACKGROUND: The historical orogenesis and associated climatic changes of mountain areas have been suggested to partly account for the occurrence of high levels of biodiversity and endemism. However, their effects on dispersal, differentiation and evolution of many groups of plants are still unknown. In this study, we examined the detailed diversification history of Primula sect. Armerina, and used biogeographic analysis and macro-evolutionary modeling to investigate a series of different questions concerning the evolution of the geographical and ecological distribution of the species in this section. RESULTS: We sequenced five chloroplast and one nuclear genes for species of Primula sect. Armerina. Neither chloroplast nor nuclear trees support the monophyly of the section. The major incongruences between the two trees occur among closely related species and may be explained by hybridization. Our dating analyses based on the chloroplast dataset suggest that this section began to diverge from its relatives around 3.55 million years ago, largely coinciding with the last major uplift of the Qinghai-Tibet Plateau (QTP). Biogeographic analysis supports the origin of the section in the Himalayan Mountains and dispersal from the Himalayas to Northeastern QTP, Western QTP and Hengduan Mountains. Furthermore, evolutionary models of ecological niches show that the two P. fasciculata clades have significantly different climatic niche optima and rates of niche evolution, indicating niche evolution under climatic changes and further providing evidence for explaining their biogeographic patterns. CONCLUSION: Our results support the hypothesis that geologic and climatic events play important roles in driving biological diversification of organisms in the QTP area. The Pliocene uplift of the QTP and following climatic changes most likely promoted both the inter- and intraspecific divergence of Primula sect. Armerina. This study also illustrates how niche evolution under climatic changes influences biogeographic patterns.

Estimating the Ontogenetic Status of an Enantiornithine Bird from the Lower Barremian of El Montsec, Central Pyrenees, Spain.

An Enantiornithes specimen from El Montsec was initially described as an immature individual based upon qualitative traits such as its relatively large orbit and overall proportions of the skull and the postcranium. In this study we re-evaluate the precise determination of the ontogenetic stage of this individual, establishing a cross-talk among taphonomic, anatomic, and morphometric data. The exceptional preservation of the specimen has allowed pondering ontogenetic influence versus preservational bias in features like the external patterns of bone surfaces, instead of being aprioristically considered due to taphonomic alterations only. The rough texture of the periosteal bone associated with pores in the distal, proximal and mid-shaft areas of the humeral shaft, indicates a subadult stage when compared with long bones of modern birds. Forelimb proportions of embryo and juvenile Enanthiornithes are equivalent to those of adult individuals of other taxa within this clade, though this is not a reliable criterion for establishing a precise ontogenetic stage. The El Montsec specimen may be attributed a close adulthood, yet only if growth regimes in Enantiornithes are considered equivalent to those in Neornithes birds.

Community structure and water quality in Mediterranean streams of a Natural Park (Sant Llorenç del Munt, NE Spain)

Se estudian las comunidades de macroinvertebrados de los ríos del Parque Natural de Sant Llorenç del Munt i la Serra de l'Obac (Barcelona, NE España) y su relación con las condiciones de flujo de los ríos. Hasta 78 localidades se visitaron en dos ocasiones (invierno y verano de 1996) y en 26 de ellas se tomaron muestras de macroinvertebrados. Mientras en invierno el 63 % de los kilÛmetros investigados tenía flujo continuo y menos del 1% estaba seco, en verano solo el 26% tenía flujo continuo, un 20% estaba totalmente seco y el resto presentaba pozas en su lecho. A pesar de ello el número de familias de macroinvertebrados fue de 54 en invierno y 94 en verano, siendo dominantes en este último caso los heterópteros, coleópteros, odonatos y dípteros, mientras que en invierno los tricópteros y plecópteros eran más diversos. En general, la comunidad presentó una estrategia trófica recolectora aunque la proporción de los ramoneadores y depredadores aumentó en verano. El estudio de las comunidades mediante el análisis de su abundancia en los dos perÌodos, mostró que las variables temporales (flujo, temperatura) o las relacionadas con el incremento de la producciÛn primaria (oxÌgeno, pH) explicaban la mayor parte de la variabilidad con los elementos mas reófilos propios de invierno y los leníticos de verano, mientras que otros factores fisicoquímicos no eran relevantes. Calculado el índice biológico BMWP' se demostró que los valores en verano eran superiores o similares a los de invierno lo que se explica por la mayor diversidad aunque la calificaciÛn individual de cada una de las familias encontradas en verano fuera menor que las halladas en invierno.

Mesoscale numerical analysis of the historical November 1982 heavy precipitation event over Andorra (Eastern Pyrenees)

From 6 to 8 November 1982 one of the most catastrophic flash-flood events was recorded in the Eastern Pyrenees affecting Andorra and also France and Spain with rainfall accumulations exceeding 400 mm in 24 h, 44 fatalities and widespread damage. This paper aims to exhaustively document this heavy precipitation event and examines mesoscale simulations performed by the French Meso-NH non-hydrostatic atmospheric model. Large-scale simulations show the slow-evolving synoptic environment favourable for the development of a deep Atlantic cyclone which induced a strong southerly flow over the Eastern Pyrenees. From the evolution of the synoptic pattern four distinct phases have been identified during the event. The mesoscale analysis presents the second and the third phase as the most intense in terms of rainfall accumulations and highlights the interaction of the moist and conditionally unstable flows with the mountains. The presence of a SW low level jet (30 m s-1) around 1500 m also had a crucial role on focusing the precipitation over the exposed south slopes of the Eastern Pyrenees. Backward trajectories based on Eulerian on-line passive tracers indicate that the orographic uplift was the main forcing mechanism which triggered and maintained the precipitating systems more than 30 h over the Pyrenees. The moisture of the feeding flow mainly came from the Atlantic Ocean (7-9 g kg-1) and the role of the Mediterranean as a local moisture source was very limited (2-3 g kg-1) due to the high initial water vapour content of the parcels and the rapid passage over the basin along the Spanish Mediterranean coast (less than 12 h).

Las hornblenditas,gabros y dioritas del Macizo del Montnegre (Barcelona, Cordilleras costero catalanas)

En relación con los granitos tardihercínicos calcoalcalinos del Macizo del Montnegre aparecen rocas ultrabásicas-básicas a intermedias. Las más básicas son hornblenditas olivínicas, con espinela, clinopiroxeno, ortopiroxeno, anfíboles de varios tipos y flogopita. Hay además melagabros-gabros hornbléndicos y leucogabros, estos últimos con anfíbol minoritario. Las rocas más ácidas son cuarzodioritas y cuarzomonzodioritas biotítico-hornbléndicas, con cristales relictos de clinopiroxeno y ortopiroxeno. El contenido máximo en anortita de la plagioclasa en las dioritas es tan alto como en los gabros. Toda la asociación muestra una evolución típicamente calcoalcalina, pero las hornblenditas y algunos gabros podrían haberse formado a partir de diferentes tipos de magmas básicos. Las cuarzodioritas serían rocas híbridas, con participación de un magma más básico y otro más ácido. El primero estaría saturado en clinopiroxeno y ortopiroxeno, además de en plagioclasa de composición bitownita, y el otro magma sería más ácido.

Estratigrafia del Cretacico inferior continental de la Cordillera Iberica Central

El Cretácico inferior continental de la Cordillera Ibérica Central está representado en dos subcuencas de sedimentación: las Cubetas de Aguilón y Oliete. Los estudios realizados sobre estos materiales han permitido definir nuevas unidades litoestratigráficas, cuya caracterización bioestratigráfica se realiza a partir de su contenido en carofitas. La identificación de discontinuidades significativas permiten definir tres secuencias de depósito que abarca en el intervalo temporal Valanguniense y/o Hauteriviense-Barremiense inferior y que son correlacionables con las S.D.K1.2 y/o K1.3, K1.4 YK1.5 y/o K1.6 definidas en Salas et al. (1991) y Salas y Casas (1993) para la Cuenca del Maestrazgo. Estos resultados ponen de manifiesto que el registro estratigráfico de la zona central de la Cordillera Ibérica tiene una entidad similar al de los márgenes de las Cuencas del Maestrazgo, Aliaga- Peñagolosa y Cameros.

Termobarometria opx-cpx aplicada al conocimiento de las condiciones de formacion de las rocas ultramaficas de Vivero (Lugo, Noroeste de EspaÑa)

Diversos métodos termométricos y barométricos, basados en el equilibrio ortopiroxeno-clinopiroxeno, se utilizan en la determinación de temperatura y presión de formación de rocas ultramáficas ricas en alfíbol, asociadas a granitos calcoalcalinos del mazizo de Vivero (Lugo, noroeste de España)

Primeros datos cromoestratigráficos de las series evaporíticas del triásico superior de Valencia (España)

El Triásico Superior del sector central valenciano de la Cordillera Ibérica presenta una facies germánica (Keuper) integrada por dos secuencias evaporíticas separadas por un episodio detrítico principal. Los datos preliminares obtenidos de una investigación palinológica en curso revelan una edad Karniense para todos estos materiales.

Structural control on present-day topography of a basement massif: the Central and Eastern Anti-Atlas (Morocco)

The Anti-Atlas basement massif extends South of the High Atlas, and, despite a very mild Cenozoic deformation, its altitude exceeds 1500m in large areas, reaching 3305m in Jbel Sirwa. Structural contours of the present elevation of a polygenic planation surface (the High Erosional surface) and of the base of Cretaceous and Neogene inliers have been performed to characterize the major tectonic structures. Gentle Cenozoic WSW-ENE- and N-Strending folds, of 60 to100km wavelength, reactivate Variscan structures, being the major contributors to the local topography of the Anti-Atlas. Reactivated thrusts of decakilometric to kilometric-scale and E-W trend involving the Neogene rocks exhibit a steep attitude and a small displacement, but they also produce a marked topographic expression. The resulting Cenozoic horizontal shortening along N-S sections across the Anti-Atlas is about 1%. The position of the major anticlinal hinges determines the location of the fluvial divides of the Warzazat basin and the Anti-Atlas, and a structural depression on one of these hinges (Jbel Saghro anticline) allowed the formerly endorheic Warzazat basin to drain southwards. The first Cenozoic structures generating local topography are of pre-mid Miocene age (postdated by 6.7Ma volcanic rocks at the Jbel Saghro), whereas the youngest thrust movements postdate the Pliocene sedimentary and volcanic rocks (involving 2.1Ma volcanic rocks at Jbel Sirwa). In addition to these features, the mean elevation of the Anti-Atlas at the regional scale is also the result of a mantle thermal anomaly reported in previous works for the entire Atlas system.

Análisis microestructural de los afloramientos en las calizas cretácicas del macizo del Garraf (prov. de Barcelona, España)

LA aplicación del modelo de la 'deformación discontinua' propuesto pr F. ARTHAUD y M. MATTAUER al análisis de la tectónica frágil de dos canteras del macizo de Garraf (prov. de Barceona, España), permite diferenciar las microestructuras pertenecientes a dos deformaciones distintas ('compresión' y 'distendión·) y caracterizar las direcciones principales de deformación para cada una de ellas.

Cenozoic evolution of eastern Iberia: structural data and dynamic model.

Iberia underwent intraplate deformation during the Mesozoic and Cenozoic. In eastem Ibena, compression took place during the Palaeogene and early Miocene, giving rise to the Iberian Chain, and extension started during the early Miocene in the coastal areas and the Valencia trough; during early Miocene compression continued in the western Iberian Chain whereas extension had started in the eastern Iberian Chain. From the kinematic data obtained from the major compressional and extensional structures formed dunng the Cenozoic, a simple dynamic model using Bott's (1959) formula is presented. The results show that both extension and compression may have been produced assuming a main horizontal stress-axis approximately N-S, in a similar direction that the convergence between Europe, Ibena and Afnca dunng the Cenozoic.

Primeros datos cromoestratigráficos de las series evaporíticas del triásico superior de Valencia (España)

El Triásico Superior del sector central valenciano de la Cordillera Ibérica presenta una facies germánica (Keuper) integrada por dos secuencias evaporíticas separadas por un episodio detrítico principal. Los datos preliminares obtenidos de una investigación palinológica en curso revelan una edad Karniense para todos estos materiales.