949 resultados para Semi-arid region


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While much is known about the factors that control each component of the terrestrial nitrogen (N) cycle, it is less clear how these factors affect total N availability, the sum of organic and inorganic forms potentially available to microorganisms and plants. This is particularly true for N-poor ecosystems such as drylands, which are highly sensitive to climate change and desertification processes that can lead to the loss of soil nutrients such as N. We evaluated how different climatic, abiotic, plant and nutrient related factors correlate with N availability in semiarid Stipa tenacissima grasslands along a broad aridity gradient from Spain to Tunisia. Aridity had the strongest relationship with N availability, suggesting the importance of abiotic controls on the N cycle in drylands. Aridity appeared to modulate the effects of pH, plant cover and organic C (OC) on N availability. Our results suggest that N transformation rates, which are largely driven by variations in soil moisture, are not the direct drivers of N availability in the studied grasslands. Rather, the strong relationship between aridity and N availability could be driven by indirect effects that operate over long time scales (decades to millennia), including both biotic (e.g. plant cover) and abiotic (e.g. soil OC and pH). If these factors are in fact more important than short-term effects of precipitation on N transformation rates, then we might expect to observe a lagged decrease in N availability in response to increasing aridity. Nevertheless, our results suggest that the increase in aridity predicted with ongoing climate change will reduce N availability in the Mediterranean basin, impacting plant nutrient uptake and net primary production in semiarid grasslands throughout this region.

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Sudamérica es una de las zonas con mayor cantidad de bosque seco tropical a nivel mundial. No obstante, estos bosques han sido poco conocidos y la mayoría de estudios han estado orientados hacia los bosques húmedos tropicales. Los bosques secos se han reducido drásticamente y siguen muy amenazados, corriendo el riesgo de desaparecer en los próximos años. Por ello, es fundamental, generar investigación aplicada para la conservación inmediata de los ecosistemas secos tropicales. En Ecuador, la situación no es diferente y las zonas secas catalogadas como biodiversas están en constante amenaza. Los pocos estudios realizados en Ecuador sobre zonas secas, han permitido mejorar nuestro conocimiento referente a la diversidad y abundancia de las especies, relaciones planta-planta y síndromes de dispersión. No existen estudios sobre caracteres morfológicos en frutos y semillas de las especies leñosas de bosque seco. Sin embargo, nuestra comprensión de la dinámica y estructura de las comunidades ecológicas de zonas secas poco estudiadas, puede mejorar rápidamente mediante el estudio y enfoque de rasgos morfofisiológicos funcionales. El objetivo general del presente estudio fue aportar al conocimiento de la ecología y biología de semillas de zonas secas tropicales mediante el análisis y evaluación de rasgos morfofisiológicos de frutos y semillas de una comunidad de especies leñosas. El estudio se realizó en una zona de bosque y matorral seco, ubicados al sur occidente del Ecuador, a una altitud comprendida entre los 250 a 1 200 m s.n.m. caracterizada por una marcada estacionalidad ambiental, con lluvias desde diciembre a abril y una estación seca de mayo a noviembre. Precipitación media anual de 500 mm con una temperatura media anual de 20° a 26 °C. La zona de estudio forma parte de la región Tumbesina compartida entre el sur del Ecuador y el norte del Perú con gran diversidad de especies vegetales endémicas. Para el estudio se colectaron frutos con semillas maduras previamente a su dispersión de entre ocho y diez individuos de 80 especies entre árboles y arbustos más representativos de los bosques secos ecuatorianos. De los frutos colectados se utilizó una muestra al azar de 50 frutos y semillas por especie para los diferentes análisis. Se midió y evaluó 18 rasgos morfológicos y fisiológicos cuantitativos y cualitativos de frutos, semillas y de la especie. Se realizaron diferentes análisis de asociación y correlación entre los rasgos evaluados, con cinco variables ambientales registradas de las 109 parcelas establecidas en el área de estudio, además analizamos el tipo de dormición y comparamos la respuesta germinativa a la deshidratación relacionada con dos comunidades secas, matorral y bosque seco. Los resultados mostraron que las especies presentan gran heterogeneidad en rasgos continuos de las semillas. La variabilidad fue más evidente en rasgos como tamaño, volumen, masa y número de semillas por fruto. Sin embargo, una alta proporción de las especies tiende a producir una semilla por fruto. Además, la mayoría de las especies de bosque seco se caracterizan por no poseer algún tipo de apéndices o areola en sus semillas, forma ovalada y sin endospermo. La reserva nutritiva de las semillas se encuentra especialmente en los cotiledones de los embriones. Se encontraron seis tipos diferentes de embriones y la mayoría de las especies presentó embriones gruesos e invertidos. La dispersión de semillas está dominada por zoocoria en un 38 %, con relación a anemocoria (22 %) y autocoria (19 %). Sin embargo, encontramos que el 70 % de las especies posee frutos secos. Los análisis de dormición en las semillas de bosque seco, mostraron que el 60 % de las especies de bosque seco presentaron semillas con algún tipo de latencia, menor a la encontrada en especies de bosque deciduo tropical y sabanas, sin embargo, la dormición de las especies de bosque seco fue mayor al porcentaje de especies con dormición de bosque semiperenne y selva lluviosa tropical. La dormición física constituyó el 35 % de las especies de bosque seco, seguido del 12 % con dormición fisiológica, mientras que solamente una especie tuvo dormición morfológica. Encontramos que la dormición de las semillas de las especies en estudio se relaciona significativamente con el tipo y función del embrión y con el endospermo. Existieron relaciones significativas entre los rasgos morfológicos de los frutos, semillas, embriones y atributos de los individuos de 46 especies, aunque en algunos casos con coeficientes de correlación bajos. Hubo pocas relaciones entre los rasgo morfológicos de las semillas con las variables ambientales registradas. Solamente el tipo de testa y la presencia de apéndices en las semillas mostraron relación con el pH y la temperatura media del suelo. No obstante usando el modelo fouth corner-RLQ, no se encontraron asociaciones claras ni significativas entre rasgos morfológicos de semillas y frutos con variables ambientales. Al medir el efecto de la deshidratación en las semillas de los dos hábitats secos tropicales: bosque y matorral seco, los resultados determinaron que tanto las semillas de las especies leñosas de ambientes más áridos (matorral seco) están en gran medida pre-adaptadas a la desecación que las especies de ambientes menos áridos (bosque seco). Los tratamientos de deshidratación ejercieron un efecto negativo en los porcentajes de germinación en todas las especies, excepto para C. platanifolia. Los resultados más sorprendentes se registraron para Senna alata que mostró germinación extremadamente baja o incluso sin germinación a contenidos de humedad de la semillas de 0,10 g H2O g de peso seco. Las curvas de germinación difirieron significativamente entre los tratamientos de deshidratación en cada especie. Aportar al conocimiento la fisiología de la deshidratación y los límites de tolerancia de las semillas de bosque y matorral seco ayudará a entender mejor el papel de este rasgo en la ecología de las semillas y dinámica de las comunidades áridas tropicales. El estudio demostró, que la adaptación ecológica de las semillas de las especies leñosas de bosque seco a factores ambientales extremos, puede verse reflejada en una red de interacciones y correlaciones complejas entre los propios rasgos morfológicos y fisiológicos continuos y cuantitativos, sobre todo en rasgos internos de las semillas, quienes ejercerían una mayor influencia en toda la red de interacciones. Si bien, los rasgos de las semillas no mostraron fuertes relaciones con las variables ambientales, posiblemente las asociaciones presentes entre rasgos morfológicos pudiesen predecir en cambio interacciones entre especies y comportamientos y procesos relacionados con la tolerancia a la deshidratación y dormición de las semillas. ABSTRACT South America is one of the areas with the largest number of tropical dry forest in the world. However, these forests have been poorly understood and most studies have been directed to tropical rainforests. Dry forests have been drastically reduced and are very threatened, risking desaparecerer in the next years. It is therefore essential, generate applied research for conservation of tropical dry ecosystems. In Ecuador the situation is no different and dry areas classified as biodiverse are under constant threat. The few studies made in Ecuador on drylands have improved our knowledge concerning the diversity and abundance of species, plant-plant relationships and dispersion syndromes. Morphological studies on fruits and seeds of woody dry forest species do not exist. However, our understanding of the dynamics and structure of ecological communities dryland little studied, may improve quickly through the study and functional approach morphophysiological traits. The overall objective of this study was to contribute to the knowledge of the ecology and biology of tropical dry seeds through analysis and evaluation of morphophysiological traits of fruits and seeds of a community of woody species. The study was conducted in an area of dry scrub forest, located at the southwest of Ecuador, at an altitude between 250 to 1200 m asl. Environmental characterized by a marked seasonality, with rainfall from December to April and a dry season from May to November. Annual rainfall of 500 mm with an average annual temperature of 20° to 26 °C. The study area is part of the shared Tumbesina region between southern Ecuador and northern Peru with a great diversity of endemic plant species. For the study, we collected fruit and seed madure of eight and ten individuos of 80 species of trees and shrub most representated of the Ecuador dry forest. We selected a sample of 50 fruits and seeds for different analysis. We measure and evaluate 18 morphological and physiological traits of fruits, seeds and species. We perform analysis and correlation between traits associated with five environmental variables taken from the 109 plots established in the study area also analyze and compare the germination response to dehydration related to two dry communities, scrub and dry forest. The results showed that the species have great heterogeneity in continuous seed traits. Variability was more evident in features such as size, volume, mass, and number of seeds per fruit. However, a high proportion of species tends to produce a seed per fruit. In addition, most of the species of dry forest is characterized by not having some sort of ppendices or areola in its seeds, oval form and without endosperm. The nutrient reserves of seeds are especially in the cotyledons of the embryos. Six different embryos were found and most of the species presented thick and inverted embryos. Seed dispersal zoochory is dominated by 38 %, relative to anemochory (22 %) and autochory (19 %). However, we found that 70 % of the species has dried fruits. The analysis of dormancy from tropical dry forest, showed that 60 % of species showed seed dormancy, down from species found in tropical deciduous forest and savanna, however dormancy dry forest species was higher than the percentage of forest species dormancy semi-evergreen and tropical rain forest. Physical dormancy corresponds to 35 % of species, followed by 12 % with physiological dormancy, while only one species had morphological dormancy. We found that dormancy of the seeds was significantly related to the type and function of the embryo and the endospemo. There were significant relationships between morphological traits of fruits, seeds, embryos and attributes of individuals of 46 species, although in some cases with low correlation coefficients. There was little relationship between the morphologic traits of the seeds with the registered environmental variables. Only the type of tesla and the presence of appendages on the seeds showed relation to pH and the mean soil temperature. However, using the fourth corner-RLQ model, neither clear nor significant between morphological traits of seeds and fruits associations with environmental variables were found. The effect of dehydration on seeds of two tropical dry forest habitats was evident in dry scrub. The results determined that both the seeds of woody species forest and dry scrub are pre-adapted to drier conditions. Dehydration treatments exerted a negative effect on germination percentage in all species, except for C. platanifolia. However, all species germinated in treatments of extreme dryness, but in low percentages. The most striking results were recorded for Senna alata showed no germination when its moisture content was 0.10 g H2O g dry weight. Germination curves differ significantly between the treatments of dehydration in each species. Contribute to the knowledge of physiology and dehydration tolerance limits seeds dry scrub forest and help you better understand the role of this trait in seed ecology and dynamics of tropical arid communities. The study showed that the ecological adaptation of seeds of woody species of dry forest to extreme environmental factors may be reflected in a complex web of interactions and correlations between morphological and physiological traits continuous and quantitative themselves, especially in internal seed traits, who exerted a major influence on the entire network of interactions. While the seed traits showed strong relationships with environmental variables possibly present associations between morphological traits could predict interactions between species and change behaviors related to desiccation tolerance and seed dormancy processes.

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Natural ecosystems contain many individuals and species interacting with each other and with their abiotic environment. Such systems can be expected to exhibit complex dynamics in which small perturbations can be amplified to cause large changes. Here, we document the reorganization of an arid ecosystem that has occurred since the late 1970s. The density of woody shrubs increased 3-fold. Several previously common animal species went locally extinct, while other previously rare species increased. While these changes are symptomatic of desertification, they were not caused by livestock grazing or drought, the principal causes of historical desertification. The changes apparently were caused by a shift in regional climate: since 1977 winter precipitation throughout the region was substantially higher than average for this century. These changes illustrate the kinds of large, unexpected responses of complex natural ecosystems that can occur in response to both natural perturbations and human activities.

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The aim of the present study is to identify and evaluate the relationship between Woodpigeon (Columba palumbus, Linnaeus, 1758) density and different environmental gradients (thermotype, ombrotype, continentality and latitudinal), land use and landscape structure, using geographic information systems and multivariate modelling. Transects (n = 396) were developed to estimate the density of Woodpigeon in the Marina Baja (Alicante, Spain) from 2006 to 2008. The highestdensity for Woodpigeon was in September-October (1.28birds/10ha) and the lowest inFebruary-March (0.34birds/10ha). Moreover, there were more Woodpigeons in areas with a mesomediterranean thermotypethan in thermomediterranean or supramediterranean ones. There was greater densityinthe intermediate zones compared to thecoast and interior. The natural or cultural landscape had the highest Woodpigeon density (1.53birds/10ha), with both denseand clear pine forest values standing out. Therefore, it is very important to conserve these traditional landscapes with adequate management strategies in order to maintain, resident and transient Woodpigeon populations. These natural areas are open places where the Woodpigeons find food and detect the presence ofpredators. Thus, this study will enable more precise knowledge of the ecological factors (habitat variables) that intervene in the distribution of Woodpigeon populations and their density.

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Binder's numbering: v.1 and 2; v.3 and 4.

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In cold-chamber high-pressure die castings (HPDC), the microstructure consists of coarse externally solidified crystals (ESCs) that are commonly observed in the central region of cross sections. In the present work, controlled laboratory scale casting experiments have been conducted with particular emphasis on the flow and solidification conditions. An A356 aluminum alloy was used to produce castings by pouring semi-solid metal through a steel die. Microstructures similar to those encountered in HPDC have been produced and the resulting microstructure is found to depend on the melt and die temperature: (1) the fraction of ESCs determines the extent of migration to the central region; (2) a maximum packing determines the area fraction of ESCs in the center; and (3) the die temperature affects the position of the ESCs-a higher die temperature can induce a displaced ESC distribution. It is found that the n-figration of crystals to the central region requires a flow, which is constrained at all melt/die interfaces. Furthermore, potential lift mechanisms are discussed. An assessment of the Saffman lift force on individual particles shows it has no significant effect on the migration of ESCs.

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A high resolution marine pollen record from site GeoB1023, west of the northern Namib desert provides data on vegetation and climate change for the last 21 ka at an average resolution of 185 y. Pollen and spores are mainly delivered to the site by the Cunene river and by surface and mid-tropospheric wind systems. The main pollen source areas are located between 13°S and 21°S, which includes the northern Namib desert and semi-desert, the Angola-northern Namibian highland, and the north-western Kalahari. The pollen spectra reflect environmental changes in the region. The last glacial maximum (LGM) was characterised by colder and more arid conditions than at present, when a vegetation with temperate elements such as Asteroideae, Ericaceae, and Restionaceae grew north of 21°S. At 17.5 ka cal. B.P., an amelioration both in temperature and humidity terminated the LGM but, in the northern Kalahari, mean annual rainfall in the interval 17.5-14.4 ka cal. B.P. was probably 100-150 mm lower than at present (400-500 mm/y). The Late-glacial to early Holocene transition includes two arid periods, i.e. 14.4-12.5 and 10.9-9.3 ka cal. B.P. The last part of the former period may be correlated with the Younger Dryas. The warmest and most humid period in the Holocene occurred between 6.3 and 4.8 ka cal. B.P. During the last 2000 years, human impact, as reflected by indications of deforestation, enhanced burning and overgrazing, progressively intensified.

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Turbulence and internal waves are probably important in generating layered structures in frontal region of marine environments (e.g. near river plumes outflow into the sea). Here we investigate the role of normal modes of internal waves in generation of layered structure in a part of Persian Gulf where river plume inters and in some laboratory experiments. The model prediction and observations show that layers so formed have a thickness of about 2m based on salinity variations with depth, but layers (about 5m) based on horizontal velocity profiles. Laboratory experiments with a plume outflow in a Filling Box profile also generate normal mode layered structure with 21H=0.5 (where A is layer thickness and H is the plume depth). In these experiments as Re of the flow is smaller than the Re of field flow. The normal modes are substantially dissipated with depth. Typical values of dissipation factor is about 0(100). This factor for field observation is 0(10) which is still substantial. Qualitative comparison between layered structure in field and laboratory is good. It should be emphasized that field observation is for semi-enclosed seas but the laboratory experiments are for enclosed region. Hence some of the discrepancies in the results of two cases are inevitable. Layered structures in marine environments are also produced by double diffusive convection. In this region this should be studied separately.

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In this work, we study the Zeeman splitting effects in the parallel magnetic field versus temperature phase diagram of two-dimensional superconductors with one graphene-like band and the orbital effects of perpendicular magnetic fields in isotropic two-dimensional semi-metallic superconductors. We show that when parallel magnetic fields are applied to graphene and as the intraband interaction decreases to a critical value, the width of the metastability region present in the phase diagram decreases, vanishing completely at that critical value. In the case of two-band superconductors with one graphene-like band, a new critical interaction, associated primarily with the graphene-like band, is required in order for a second metastability region to be present in the phase diagram. For intermediate values of this interaction, a low-temperature first-order transition line bifurcates at an intermediate temperature into a first-order transition between superconducting phases and a second-order transition line between the normal and the superconducting states. In our study on the upper critical fields in generic semi-metallic superconductors, we find that the pair propagator decays faster than that of a superconductor with a metallic band. As result, the zero field band gap equation does not have solution for weak intraband interactions, meaning that there is a critical intraband interaction value in order for a superconducting phase to be present in semi-metallic superconductors. Finally, we show that the out-of-plane critical magnetic field versus temperature phase diagram displays a positive curvature, contrasting with the parabolic-like behaviour typical of metallic superconductors.

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The influence of particles recycling on the geochemistry of sediments in a large tropical dam lake in the Amazonian region, Brazil. Article in Journal of South American Earth Sciences 72 · December 2016 DOI: 10.1016/j.jsames.2016.09.012 1st Rita Fonseca 16.85 · Universidade de Évora 2nd Catarina Pinho 3rd Manuela Oliveira 22.6 · Universidade de Évora Abstract As a result of over-erosion of soils, the fine particles, which contain the majority of nutrients, are easily washed away from soils, which become deficient in a host of components, accumulating in lakes. On one hand, the accumulation of nutrients-rich sediments are a problem, as they affect the quality of the overlying water and decrease the water storage capacity of the system; on the other hand, sediments may constitute an important resource, as they are often extremely rich in organic and inorganic nutrients in readily available forms. In the framework of an extensive work on the use of rock related materials to enhance the fertility of impoverish soils, this study aimed to evaluate the role on the nutrients cycle, of particles recycling processes from the watershed to the bottom of a large dam reservoir, at a wet tropical region under high weathering conditions. The study focus on the mineralogical transformations that clay particles undergo from the soils of the drainage basin to their final deposition within the reservoir and their influence in terms of the geochemical characteristics of sediments. We studied the bottom sediments that accumulate in two distinct seasonal periods in Tucuruí reservoir, located in the Amazonian Basin, Brazil, and soils from its drainage basin. The surface layers of sediments in twenty sampling points with variable depths, are representative of the different morphological sections of the reservoir. Nineteen soil samples, representing the main soil classes, were collected near the margins of the reservoir. Sediments and soils were subjected to the same array of physical, mineralogical and geochemical analyses: (1) texture, (2) characterization and semi-quantification of the clay fraction mineralogy and (3) geochemical analysis of the total concentration of major elements, organic compounds (organic C and nitrogen), soluble fractions of nutrients (P and K), exchangeable fractions (cation exchange capacity, exchangeable bases and acidity) and pH(H2O).

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Scorpion stings account for most envenomations by venomous animals in Brazil. A retrospective study (1994-2011) of the clinical consequences of Tityus scorpion stings in 1327 patients treated at a university hospital in Campinas, southeastern Brazil, is reported. The clinical classification, based on outcome, was: dry sting (no envenoming), class I (only local manifestations), class II (systemic manifestations), class III (life-threatening manifestations, such as shock and/or cardiac failure requiring inotropic/vasopressor agents, and/or respiratory failure), and fatal. The median patient age was 27 years (interquartile interval = 15-42 years). Scorpions were brought for identification in 47.2% of cases (Tityus bahiensis 27.7%; Tityus serrulatus 19.5%). Sting severity was classified and each accounted for the following percentage of cases: dry stings - 3.4%, class I - 79.6%, class II - 15.1%, class III - 1.8% and fatal - 0.1%. Pain was the primary local manifestation (95.5%). Systemic manifestations such as vomiting, agitation, sweating, dyspnea, bradycardia, tachycardia, tachypnea, somnolence/lethargy, cutaneous paleness, hypothermia and hypotension were detected in class II or class III + fatal groups, but were significantly more frequent in the latter group. Class III and fatal cases occurred only in children <15 years old, with scorpions being identified in 13/25 cases (T. serrulatus, n = 12; T. bahiensis, n = 1). Laboratory blood abnormalities (hyperglycemia, hypokalemia, leukocytosis, elevations in serum total CK, CK-MB and troponin T, bicarbonate consumption and an increase in base deficit and blood lactate), electrocardiographic changes (ST segment) and echocardiographic alterations (ventricular ejected fraction <54%) were frequently detected in class III patients. Seventeen patients developed pulmonary edema, 16 had cardiac failure and seven had cardiogenic shock. These results indicate that most scorpion stings involved only local manifestations, mainly pain; the greatest severity was associated with stings by T. serrulatus and in children <15 years old.

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Giardia duodenalis is a flagellate protozoan that parasitizes humans and several other mammals. Protozoan contamination has been regularly documented at important environmental sites, although most of these studies were performed at the species level. There is a lack of studies that correlate environmental contamination and clinical infections in the same region. The aim of this study is to evaluate the genetic diversity of a set of clinical and environmental samples and to use the obtained data to characterize the genetic profile of the distribution of G. duodenalis and the potential for zoonotic transmission in a metropolitan region of Brazil. The genetic assemblages and subtypes of G. duodenalis isolates obtained from hospitals, a veterinary clinic, a day-care center and important environmental sites were determined via multilocus sequence-based genotyping using three unlinked gene loci. Cysts of Giardia were detected at all of the environmental sites. Mixed assemblages were detected in 25% of the total samples, and an elevated number of haplotypes was identified. The main haplotypes were shared among the groups, and new subtypes were identified at all loci. Ten multilocus genotypes were identified: 7 for assemblage A and 3 for assemblage B. There is persistent G. duodenalis contamination at important environmental sites in the city. The identified mixed assemblages likely represent mixed infections, suggesting high endemicity of Giardia in these hosts. Most Giardia isolates obtained in this study displayed zoonotic potential. The high degree of genetic diversity in the isolates obtained from both clinical and environmental samples suggests that multiple sources of infection are likely responsible for the detected contamination events. The finding that many multilocus genotypes (MLGs) and haplotypes are shared by different groups suggests that these sources of infection may be related and indicates that there is a notable risk of human infection caused by Giardia in this region.

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The aim of this study was to estimate barite mortar attenuation curves using X-ray spectra weighted by a workload distribution. A semi-empirical model was used for the evaluation of transmission properties of this material. Since ambient dose equivalent, H(⁎)(10), is the radiation quantity adopted by IAEA for dose assessment, the variation of the H(⁎)(10) as a function of barite mortar thickness was calculated using primary experimental spectra. A CdTe detector was used for the measurement of these spectra. The resulting spectra were adopted for estimating the optimized thickness of protective barrier needed for shielding an area in an X-ray imaging facility.

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Primary X-ray spectra were measured in the range of 80-150kV in order to validate a computer program based on a semiempirical model. The ratio between the characteristic and total air Kerma was considered to compare computed results and experimental data. Results show that the experimental spectra have higher first HVL and mean energy than the calculated ones. The ratios between the characteristic and total air Kerma for calculated spectra are in good agreement with experimental results for all filtrations used.