Uppermost Limit, Extent, and Fluctuations of the Timberline and Treeline Ecocline in the Swiss Central Alps during the past 11,500 Years

Pollen and macrofossils were analyzed at two sites above today's treeline (or tree limit) in the Swiss Central Alps (Gouillé Loéré, 2503 m a.s.l., and Lengi Egga, 2557 m a.s.l.) to test two contrasting hypotheses about the natural formation of timberline (the upper limit of closed forest) in the Alps. Our results revealed that Pinus cembra--Larix decidua forests near timberline were rather closed between 9000 and 2500 B.C. (9600-4000 ¹⁴C yr BP), when timberline fluctuations occurred within a belt 100-150 m above today's tree limit. The treeline ecocline above timberline was characterized by the mixed occurrence of tree, shrub, dwarf-shrub, and herbaceous species, but it did not encompass more than 100-150 altitudinal meters. The uppermost limit reached by timberline and treeline during the Holocene was ca. 2420 and 2530 m, respectively, i.e., about 120 to 180 m higher than today. Between 3500 and 2500 B.C. (4700-4000 ¹⁴C yr BP) timberline progressively sank by about 300 m, while treeline was lowered only ca. 100 m. This change led to an enlargement of the treeline-ecocline belt (by ca. 300 m) after 2500 B.C. (4000 ¹⁴C yr BP). Above the treeline ecocline, natural meadows dominated by dwarf shrubs (e.g., Salix herbacea) and herbaceous species (e.g., Helianthemum, Taraxacum, Potentialla, Leontodon t., Cerastium alpinum t., Cirsium spinosissimum, Silene exscapa t., and Saxifraga stellaris) have been present since at least 11,000 cal yr ago. In these meadows tree and tall shrub species (>0.5 m) never played a major role. These results support the conventional hypothesis of a narrow ecocline with rather sharp upper timberline and treeline boundaries and imply that today's treeless alpine communities in the Alps are close to a natural stage. Pollen (percentages and influx), stomata, and charcoal data may be useful for determining whether or not a site was treeless. Nevertheless, a reliable and detailed record of past local vegetation near and above timberline is best achieved through the inclusion of macrofossil analysis.

Morphometric redefinition of the component chronospecies of the Globorotalia conoidea - G. inflata lineage in DSDP Hole 29-284

Phylo-zonations (or lineage-zonations) are based upon morphological changes within individual evolutionary lineages. These zonations, although potentially of use for stratigraphic subdivision and correlation, often suffer from a lack of quantitative exactness in the definitions of chronospecies. Thus exact reproducibility is hindered for stratigraphic determinations. The potential of morphometrically defined phylo-zonations is demonstrated on a temperate South Pacific Late Cenozoic lineage of planktonic foraminifera (Globorotalia conoidea through intermediate forms to Globorotalia inflata in DSDP Site 284) exhibiting phyletic gradualism. Our sampling interval is about 0.1 m.y. during the last 8 m.y. Changes in the number of chambers in the final whorl, test conicalness, percentage of keeled forms, and test roundness or inflatedness, are used to quantitatively define the following five chronospecies: G. conoidea (Late Miocene; 6.1->8.3 m.y.), G. conomiozea (latest Miocene ; 5.3-6.1 m.y.), G. puncticulata sphericomiozea (earliest Pliocene; 4.5-5.3 m.y.), G. puncticulata puncticulata (Early-Middle Pliocene; 2.9-4.5 m.y.), and G. inflata (Late Pliocene-Quaternary; 0-2.9 m.y.). This phylo-zonation is directly applicable to temperate cool subtropical Southern Hemisphere areas where the evolution took place (Kennett, 1967, 1973; Scott, 1979). It is still not known if the lineage occurs elsewhere; thus the applicability of the phylo-zonation over broader areas is still uncertain. Trends in general size and aperture shape seem to be climatically controlled, and thus may be only of local stratigraphic utility. The practical applications of morphometric phylo-zonation for stratigraphy is to a large extent dependent upon the amount of time and effort required to statistically define the trends. Experiments with large numbers of subsamples from this lineage demonstrate that accurate stratigraphic determinations are possible from measurements on only 15 specimens per sample, except for those very close to chronospecies boundaries.

Synchroneity of Pliocene planktonic foraminiferal datums in the North Atlantic

Leg 94 of the Deep Sea Drilling Project has provided a unique set of paleomagnetically dated cores, taken along a N-S transect in the North Atlantic. High deposition rates in the sediments, combined with the palaeomagnetic ages, have enabled existing planktonic foraminiferal zonations to be tested and a new zonation for the mid- to high latitudes to be erected. The PL zonation of Berggren (1973, 1977) is shown to be adequate as far north as 41°N, although both the LAD's of Globigerina nepenthes and Globorotalia margaritae occur earlier than in tropical regions. North of 41°N these two species have very diachronous LAD's, even though they are common during their range in the northern sites. The new zonation for the mid to high latitude North Atlantic is based on the FAD of G. margaritae, FAD of G. puncticulata, LAD of G. cf. crassula, LAD of N. atlantica, FAD of G. inflata and FAD of sinistrally coiled encrusted N. pachyderma.

Chemical and isotopic compositions of hydrothermal manifestations and minerals from the Broken Spur hydrothermal field

A representative collection of hydrothermal manifestations was sampled practically from all hydrothermal mounds of the Broken Spur hydrothermal vent field with use of the Mir manned submersibles during three cruises of R/V Akademik Mstislav Keldysh. Mineral associations characteristic for different morphological types of sulfide ores from hydrothermal pipes, plates, and diffusers are assessed. Particular attention is paid to distribution of minor elements and their distribution patterns determined by mineralogical zonation. Measured isotopic composition of sulfur in sulfide minerals varies from 0.4 to 5.2 per mil that indicates their similarity with ores from the Snake Pit vent field and is related to dilution of hot ore-bearing solutions by seawater and reduction of water sulfate ions to H2S with heavy isotopic composition.

Late Miocene to Quaternary planktonic foraminifera of ODP SIte 114-704

The planktonic foraminiferal zonation of Jenkins and Srinivasan (1986), which was defined for the southwestern sector of the temperate South Pacific Ocean, was successfully extended to the temperate sequences at Site 704. The zonation is based on first and last appearances of globorotalids, principally Globoconella species, which are indigenous to temperate surface-water masses. Most of the first and last appearances at ODP Site 704 are diachronous with those in the warmer temperate to subtropical South Atlantic, North Atlantic, and South Pacific oceans. The upper Miocene, upper Pliocene, and Quaternary sequences are punctuated by frequent incursions of subantarctic and polar assemblages of planktonic foraminifers. I assume that the appearance of an assemblage dominated by sinistral Neogloboquadrina pachyderma means that the Polar Front has migrated northward, but I do not know its position north or south of the site based on this preliminary work. The upper Miocene sequence contains five incursions between 6.5 and 5 Ma and the upper Pliocene and Quaternary sequence contains 16 events since 2.47 Ma. These are minimum estimates because the number of observed events will probably increase with higher sample density and use of quantitative methods to reveal more subtle events.

Ether lipids in deep sea and swamp sediments

Methanogen ether lipids have been quantified in sediments from a Florida swamp and the Atlantic ocean. Swamp cores containing acyclic and monocyclic isopranyl ethers are clearly differentiated from deep sea sediments which also contain bicyclic compounds. A concentration maximum near the bottom of the sulfate reducing zone in deep sea sediments may reflect a biogeochemical system in which methanogenesis and sulfate reduction are coupled by the process of methane oxidation. Lipid diagenesis is evident in the deep sea sediments. Species zonation, possibly caused by oxygen sensitivity, is detected in the relative lipid abundances in swamp sediments.

(Tab. XIII.1) TOC content, carbonate content, carbonate mineralogy, percentage of total carbonate, aragonite, low magnesium calcium, and high magnesium calcium, and grain-size contents from samples from the Golfe d'Arguin and from the Western Sahara

Modern carbonate sedimentation takes place on the northern Mauritanian shelf (20°N), where typical tropical components (e.g. hermatypic reefs, calcareous green algae) are absent. Such deposits are reminiscent of extratropical sediment in the geological record. The tropical open shelf of Mauritania is influenced by large siliciclastic dust input and upwelling, highly fertilizing the ocean, as well as strongly limiting the light penetration. In this context, temperature does not appear to be the steering factor of carbonate production. This thesis describes the depositional system of the Golfe d'Arguin off Mauritania and focuses on environmental conditions that control the depositional pattern, in particular carbonate production. The description of this modern analogue provides a tool for paleoenvironmental interpretation of ancient counterparts. The Golfe d'Arguin is a broad shallow shelf comprising extensive shoals (<10 m water depth; i.e. the Banc d'Arguin) on the inner shelf where waters warm up. The sediments collected in water depths between 4 and 600 m are characterized by mixed carbonate and siliciclastic (dust) deposits. They vary from clean coarse-grained, almost pure carbonate loose sediments to siliciclastic-dominated fine-grained sediments. The carbonate content and sediment grain size show a north-south decreasing pattern through the Golfe d'Arguin and are controlled by the hydraulic regime influenced by wind-driven surface currents, swell, and tidal currents. The carbonate grain association is heterozoan. Components include abundant molluscs, foraminifers, and worm tubes, as well as barnacles and echinoderms, elements that are also abundant in extratropical sediments. The spatial distribution of the sedimentary facies of the Golfe d'Arguin does not display a depth zonation but rather a mosaic (i.e. patchy distribution). The depth and climatic signatures of the different sedimentary facies are determined by taxonomic and ecological investigations of the carbonate-secreting biota (molluscs and foraminifers). While certain planktonic foraminifers and molluscs represent upwelling elements, other components (e.g. mollusc and benthic foraminifer taxa) demonstrate the tropical origin of the sediment. The nutrient-rich (and thus also low light-penetration) conditions are reflected in the fact that symbiotic and photosynthetic carbonate-producing organisms (e.g. hermatypic corals) are absent. The Mauritanian deposits represent an environment that is rare in the modern world but might have been more common in the geological past when global temperatures were higher. Taxonomic and ecological studies allow for distinguishing carbonate sediments formed under either tropical high-nutrient or extratropical conditions, thus improving paleoclimate reconstruction.

(Figure 4) Stratigraphic distribution of planktonic foraminifera in DSDP Hole 93-604

During Deep Sea Drilling Project Leg 93, upper Miocene through Quaternary sediments were continuously cored in Hole 604, located on the upper continental rise of the New Jersey transect (western North Atlantic). A detailed biostratigraphic study of these strata has been made using the vertical distribution of planktonic foraminifers. The Quaternary climatic zonation of Ericson and Wollin (1968) has been tentatively delineated and all the Pliocene zones and subzones (sensu Berggren, 1977) have been recognized. The rate of sedimentation was slow during most of the Pliocene but underwent a significant acceleration in the early Pleistocene. Quantitative variations in the distribution of planktonic foraminifers appear to be influenced by various factors, such as hydrodynamic winnowing resulting from the action of bottom currents and surficial thermal conditions caused by climatic changes. Both dissolution intervals and brief increases in the coarser detrital input seem, most of the time, to be correlated with indications of climatic cooling and may correspond to glacial events or cycles. This chapter delineates a precursor stage in the inception of Northern Hemisphere glaciation at 3 Ma and wide-scale Quaternary glacial-interglacial cycles. Data from a detailed study of Hole 604 are briefly compared with the main sedimentary and microfaunal features of contemporaneous series previously drilled along the east American margin in the northwestern Atlantic. One of the striking observations appears to be the intense redistribution of sediments that affected this region in Neogene-Quaternary times.

Benthic foraminifera of surface samples and sediment cores off Senegal and Gambia, West Africa

Water depth zonation of fifty nine benthonic foraminiferal species in marine sediment surfaces has been described. Some species are combined to groups which mark particular depth zones: an upper and lower shelf-fauna, an upper and lower slope fauna, and a shelf-slope fauna. Dependence on latitude could be ascertained for Textularia panamensis, and submergence effects for Hyalinea balthica.

Middle Campanian-lowermost Maastrichtian nannofossils and foraminifers of the northwestern Australian margin

The middle-late Campanian was marked by an increase in the bioprovinciality of calcareous microfossil assemblages into distinct Tethyan, Transitional, and Austral Provinces that persisted to the end of the Maastrichtian. The northwestern Australian margin belonged to the Transitional Province and the absence of key Tethyan marker species such as Radotruncana calcarata and Gansserina gansseri has led petroleum companies operating in the area to use the locally developed KCCM integrated calcareous microfossil zonation scheme. The KCCM zonation is a composite scheme comprising calcareous nannofossil (KCN), planktonic foraminiferal (KPF) and benthonic foraminiferal (KBF) zones. This paper presents the definitions and revisions of Zones KCCM8-19, from the highest occurrence (HO) of Aspidolithus parcus constrictus to the lowest occurrence (LO) of Ceratolithoides aculeus, and builds on our previous early-late Maastrichtian study. The presence of a middle-upper Campanian disconformity is confirmed by microfossil evidence from the Vulcan Sub-basin, Exmouth and Wombat plateaus, and the Southern Carnarvon Platform. In the Vulcan Sub-basin and on the Exmouth Plateau (ODP Hole 762C) the hiatus extends from slightly above the LO of common Rugoglobigerina rugosa to above the LO of Quadrum gothicum. On the Wombat Plateau (ODP Hole 761B) it spans from above the LO of Heterohelix semicostata to above the LO of Quadrum gothicum; and in the Southern Carnarvon Platform the disconformity has its longest duration from above the HO of Heterohelix semicostata to above the LO of Quadrum sissinghii. A significant revision of the events which define Zones KCCM18 and 19 was necessary owing to the observation that the LO of Ceratolithoides aculeus occurs below the HOs of Archaeoglobigerina cretacea and Stensioeina granulata incondita and the LO of common Rugoglobigerina rugosa. In the original zonation these events were considered to be coincident.

Strontium isotope ratios of planktonic foraminifera from ODP Site 119-744 on the Kerguelen Plateau, Indian Ocean (Table 1)

87Sr/86Sr ratios of well-preserved early Miocene-Oligocene planktonic foraminifers from Site 744 in the southern Indian Ocean provide the highest southern latitude Sr isotope record of this age. The isotopic data have been calibrated with the site magnetostratigraphy. 87Sr/86Sr ages were also determined using the Sr isotope-age equations of Miller et al. (1988, doi:10.1029/PA003i002p00223) and Hess et al. (1989, doi:10.1029/PA004i006p00655). There is good agreement between the calculated ages from 87Sr/86Sr measurements using these equations and those derived from magnetobiostratigraphy. In addition, these equations were useful for inference of sediment ages in intervals where the paleomagnetic record is not well resolved and the biostratigraphy is inconclusive. The Site 744 87Sr/86Sr record can be used for correlation of Antarctic and low-latitude sequences and biostratigraphical zonation of foraminifers, radiolarians, diatoms, and calcareous nannofossils. This record will assist in the development of the high southern latitude biochronology.