A biogeographically mixed, Middle Permian brachiopod fauna from the Baoshan Block, Western Yunnan, China

A small brachiopod fauna is described from the carbonate rocks of the basal Shazipo Formation of the Baoshan Block, western Yunnan, south-west China, including significant new ventral and dorsal internal morphological features of Cryptospirifer omeishanensis Huang. This fauna is regarded as Wordian (Middle Guadalupian, Middle Permian) because of the presence of Cryptospirifer omeishanensis Huang and associated fusulinids (Neoschwagerina craticulifera Zone). Palaeobiogeographically, the brachiopod fauna is of considerable interest because of its admixed nature characterized by typical warm-water Cathaysian elements intermingled with temperate Peri-Gondwanan taxa. This in turn is interpreted to indicate that the Baoshan Block may have been situated in an intermediate palaeogeographical position between Gondwanaland to the south and Cathaysia to the north during the Mid Permian and, as such, it probably furnished an important 'stepping stone' for the dispersal of Mid Permian eastern Tethyan marine invertebrate taxa (e.g. Cryptospirifer) to the western Tethys.

Possible influence of Gondwanan glaciation on low-latitude carbonate sedimentation and trans-equatorial faunal migration: the lower Permian of South China

Peculiar Early Permian palaeontological and sedimentological features are reviewed from South China, including characteristic Early Permian cold-water Gondwanan brachiopod taxa and faunas from Sichuan and Guizhou provinces, widespread rosettes and irregular aggregates of calcite prisms ('Chrysanthemum Stones') within the Qixia limestones, and lack of significant Early Permian reef buildups. The occurrences of these features are at odds with the currently widely held view that South China was located in a palaeotropical, warm-water setting throughout the Permian and hence harboured a highly diverse shallow marine biota. In this paper, I propose a working hypothesis, suggesting that influence of at least cool water masses may have intermittently occurred in South China during the Early Permian, which facilitated the formation of the cool water-influenced palaeontological and sedimentological features and promoted the interchanges of cool to cold water marine faunas between the Gondwanan and Boreal Realms. These cool water masses may have been transported to low-latitude regions as deep currents from northern and eastern shelves of Gondwanaland and upwelled along the western coast of South China as well as within the relatively deep-water basins of central South China. Prevalence of these meridional, north-directed deep cold water currents during the Early Permian may have been related to the glaciation event of Gondwanaland. An alternative and/or additional source of cooling may have also originated from strong easterly palaeoequatorial boundary currents operating within the Palaeotethys at times during the Early Permian, inducing and/or enhancing upwelling of cool to cold water masses in the eastern Palaeotethys. This latter scenario is analogous to the occasional 'La Nina' effect (opposite to the 'El Nino' effect) at the equatorial belt of the modern Pacific Ocean.

A Lopingian (Late Permian) brachiopod fauna from the Qubuerga formation at Shengmi in the Mount Qomolangma region of Southern Xizang (Tibet), China

A small fauna of 11 species belonging to 10 genera of Permian Brachiopoda from the lower part of the Qubuerga Formation outcropping near Shengmi village in the Qomolangma region of southern Xizang (Tibet) is figured and new taxa are described. New taxa are Quinquenella semiglobosa and Costatumulus shengmiensis. The fauna is most likely of Wuchiapingian (Djhulfian) age as indicated by the majority of the brachiopod species.

Late carboniferous to early permian brachiopod faunas from the Bachu and Kalpin areas, Tarim Basin, NW China

Late Carboniferous and Early Permian brachiopod faunas are described from the Xiaohaizi section of the Bachu area and the Shishichang section of the Kalpin area, the Tarim Basin, NW China. Biostratigraphic studies of brachiopods and associated microfossils indicate that the Xiaohaizi Formation is Moscovian (Late Carboniferous) and the Shishichang Formation is Kasimovian-Gzhelian (Late Carboniferous), whereas the Nanza and Kankarin Formations are Asselian to early Artinskian (Early Permian). Two new species proposed from the Nanza Formation are Kutorginella tarimensis and Phricodothyris? bachuensis.

Middle permian brachiopods from central Peninsular Malaysia - faunal affinities between Malaysia and west Cambodia

A moderately diverse Permian brachiopod fauna is described from a new rock unit, the Bera Formation, in the Bera District, central Pahang, Peninsular Malaysia. The fauna consists of 19 taxa, including 14 genera and 17 (both identified and unidentified) typically Tethyan species. The fauna appears to be correlative on the basis of brachiopods with the Neoschwagerina-Yabeina fusulinid Zones in Indochina and South China. In particular, it has strong linkage to Member C (Yabeina beds) of the Sisophon Limestone, west Cambodia. This is indicated by three of the Bera species — Urushtenoidea chaoi (Ching), Spyridiophora gubleri Termier and Termier, and Transennatia termierorum sp. nov., being shared with the Cambodian fauna. A possible early Capitanian (Middle Permian) age is proposed for the Bera brachiopod fauna.

Permian brachiopods from the Selong Xishan section, Xizang (Tibet), China. Part 2: palaeobiogeographical and palaeoecological implications, spiriferida, athyridida and terebratulida

Brachiopods from the Selong Group at the Selong Xishan section indicate a typical Peri-Gondwanan affinity, but possibly a rapid invasion of tropical/subtropical Tethyan elements into the Himalayan region at the very end of Permian. A brief analysis of taphonomy and composition of the Selong fauna indicate that Selong Xishan lay on the shallow continental shelf near shore and brachiopods suffered substantial abrasion from contemporaneous wave action. Species of Spiriferida, Athyridida and Terebratulida are herein described and revised. New taxa are Nakmusiella selongensis nov. gen. et sp. and ? Bullarina striata nov. sp.

Reworked Devonian (Givetian-Frasnian) spores from a Permian glacially-derived sequence at Bacchus Marsh, Victoria

Emphanisporites rotatus McGregor emend. McGregor 1973 is a distinctive Devonian spore with a known primary source age range in Australia spanning the upper Givetian to early Frasnian (Middle to Late Devonian). This is the first record of the species from Victoria. It occurs as a reworked element in an Early Permian assemblage belonging to the Granularisporites confluens Zone derived from glacigene diamictite in the Bacchus Marsh area. As the predominant direction of Permian ice movement recorded in the Bacchus Marsh district was south-west to north-east, it is possible that the reworked spores were transported from Antarctica.

New Strophalosiidae (Brachiopoda) from the early Permian of Argentina

New representatives of the family Strophalosiidae (Brachiopoda) are described from the Tupe Formation of the Paganzo Basin, Argentina. The genus Coronalosia Waterhouse & Gupta is reviewed and the new taxa Coronalosia argentinensis sp. nov. and Tupelosia paganzoensis gen. et sp. nov. proposed. The age of the Tupe Formation is reviewed and a middle to late Asselian (Early Permian) age is preferred.
The new genus Guadalupelosia from the mid-Permian of West Texas, USA, is also proposed.

A new genus of Rhynchonellid Brachiopod from the Lower Triassic of South China and Implications for timing the recovery of Brachiopoda after the end-Permian mass extinction

A new genus, Meishanorhynchia, is proposed based on new material from the Lower Triassic of the Meishan section, South China. It is of a late Griesbachian age based on both associated biozones (ammonoids and
bivalves) and radiometric dates of the intercalated volcanic ash beds. Comparison with both Palaeozoic and Mesozoic-Cenozoic-related genera suggests that it may represent the first radiation of progenitor brachiopods in the aftermath of the end-Permian extinction. The lowest brachiopod horizon that contains the genus is estimated to be about 250.1±0.3 Ma. This implies that the initial stage of recovery of Brachiopoda in the Early Triassic was probably about 1.3±0.3 myr after the major pulse of the end-Permian mass extinction (dated as 251.4±0.3 Ma). This is in agreement with Hallam's expectancy that biotic recovery typically begins within one million years or so of major mass extinctions, in contrast to current views on the end-Permian extinction event which propose that the recovery of most if not all biotic groups in the Early Triassic was severely delayed and only began about five million years after the end-Permian extinction.

Permophricodothyris pavlova, 1965 (Brachiopoda, Spiriferida) from the Permian of South China: its morphology, biostratigraphy and distribution

A study of both silicified and nonsilicified specimens of Permian reticularioid brachiopods from South China suggests that Permophricodothyris, a genus previously rarely reported from China, is actually very common and abundant in the Middle and especially Upper Permian of South China. This study also clarifies, for the first time, that many of the reticularioid brachiopod species previously described as Squamularia in fact belong to Permophricodothyris. The new data presented in this paper also allows a critical evaluation of Permophricodothyris in relation to its closest allies: Phricodothyris, Squamularia, Bullarina and Neophricodothyris. The revision reveals that a total of 18 Permophricodothyris species are present in the Middle and Upper Permian of South China, with only one species, P. squamularioides, having survived the Permian-Triassic mass extinction. Two species, P. grandis (Chao) and P. guangxiensis Han, Zhou & Wang, are redescribed here, providing critical new information on the morphology and taxonomy of these species.

Middle permian (Guadalupian) brachiopods from the Xiujimqinqi area, Inner Mongolia, northeast China, and their palaeobiogeographical and palaeogeographical significance

A small brachiopod fauna is described from the lower part of the Xiujimqinqi Formation of the Xiujimqinqi area in central-east Inner Mongolia, northeast China. The age of this fauna is regarded as Wordian (Middle Guadalupian, Middle Permian) by comparison with a similar brachiopod fauna from the Zhesi area of central Inner Mongolia, and by constraints from fusulinaceans associated with the Zhesi fauna. The Xiujimqinqi fauna is typical of mixed Boreal/Palaeoequatorial Middle Permian brachiopod faunas of East Asia. The mixed nature of these faunas is interpreted to have resulted from the combined effects of a middle palaeolatitudinal position, intensified plate convergence between Sino-Korea and Mongolia, and sea surface current connections with both the Arctic Sea in the north and eastern Palaeo-Tethys to the south. Possible Kaninospirifer is reported for the first time from China.

Linshuichonetes gen. nov., a new rugosochonetid (Brachiopoda) genus from the Liangshan Formation (Early Permian) in Sichuan, China, and its ecology

A new genus and species, Linshuichonetes elfinis, belonging to the Rugosochonetidae, is described from the Early Permian (Late Artinskian or early Kungurian) Liangshan Formation of the Yangtze block. The new genus is defined externally by the presence of fine, but delayed, capillation and a weak or absent median sulcus and fold and by the presence of a distinct posteromedian sinus on the ventral umbo; and internally by a lack of median, lateral and accessory septa in the dorsal interior; absence of vascular trunks in the ventral interior and the presence of distinct radiating rows of papillae in the interior of both valves, particularly an unusual clustered arrangement of papillae on the posteromedian portion of the dorsal interior. The local environment during the deposition of the Liangshan Formation appears to have been a restricted tidal flat or lagoon which experienced frequent sealevel fluctuations associated with the onset of the Yanghsingian transgression. The new species, L. elfinis, appears to have several morphological adaptations enabling successful exploitation of this environment. It was typically a very small and thin-valved species with a high surface area to volume ratio, an advantage in an oxygen restricted environment. The small size and numerous body spinules would have aided individuals to remain suspended at the top of the fine, soft substrate. It also dominated the brachiopod assemblage in the Liangshan Formation, comprising up to 94%of specimens within a bed. These factors indicate that the new species appears to be an opportunistic species.

Global distribution of permian conulariids and palaeobiogeographical implications

Previously, minimal work has been carried out on conulariids due to their rare occurrences and resultant biostratigraphical limitations. The palaeobiogeographical distribution of Permian conulariids suggests that they have a marked preference for cold to cool-water regions, that they are significant indicators for migration patterns, and that they can potentially provide information on the palaeogeographical configuration and movement of terranes. Permian conulariids are found in Australia, India, New Zealand, Pakistan, Iran, Afghanistan, Kashmir, China, Japan, Russia, Germany, Canada, United States of America, and Bolivia. The diversity of Permian conulariids is markedly higher in the polar regions than in the palaeoequatorial region.

Permian conulariid genera include Notoconularia Thomas 1969, Gondaconularia Waterhouse 1986, Cheliconularia Waterhouse 1986, Neoconularia Sugiyama 1942, Calloconularia Sinclair 1952, Diconularia Sinclair 1952, Paraconularia Sinclair 1940, Mesoconularia Boucek 1939 and Conularia Sowerby 1821. This paper describes two new species of conulariids: Diconularia meadepeakensis sp. nov. from the Phosphoria Formation (Guadalupian), Idaho, USA and Paraconularia kazanensis sp. nov. from the Sokian Horizon (?Roadian), Volga Region, Russia.

Global review of permian Tyloplecta muir-wood and cooper, 1960 (Brachiopoda): morphology, palaeobiogeographical and palaeogeographical implications

A global review of the stratigraphical and geographical distribution of Tyloplecta reveals that the genus ranges in age from Kungurian to Changhsingian (Middle to Late Permian). Tyloplecta first evolved in South China in the Kungurian (late Early Permian). The genus went through its first diversification in the Guadalupian, suffered a major extinction at the end of the Guadalupian, and re-diversified in the Wuchiapingian. T. yangtzeensis persisted into the Changhsingian as the only survivor of the genus involved in the end-Permian mass extinction. Palaeogeographically, South China is not only the centre of origin for the genus but also an area of diversification and evolution. In addition to South China, Tyloplecta has also been recorded from the Far East Russia, Japan, central Thailand, Laos, Cambodia, Qiangtang Terrane of Tibet, Salt Range, Iran, Armenia, Hungary, Yugoslavia, and Slovenia. This geographic spread suggests that Tyloplecta was primarily restricted to the Palaeotethys and is indicative of warm-water palaeoequatorial conditions. Its presence in some of the northeast Asian terranes (e.g., parts of Japan and Far East Russia) and in the Salt Range (Pakistan) and central and north Iran (part of the Cimmerian microcontinents) demonstrate that the genus invaded the middle palaeolatitudinal regions in both hemispheres during the late Middle Permian in response to increased shallow marine biotic communications between Cathaysia in the eastern Palaeotethys and southern Angaraland, and between Cathaysia and Peri-Gondwanaland. The invasion of Tyloplecta (and some other taxa) into the southern shore waters of Angaraland may be explained by assuming ocean surface current connections and close palaeogeographical proximities between the South China, Sino-Korea and Bureya blocks. In comparison, the invasion of Tyloplecta into the Peri-Gondwanaland region is more likely a result of reduced palaeogeographical distance between South China and Peri-Gondwanaland and the appearance of the Cimmerian microcontinents as migratory stepping stones.

A Guadalupian-Lopingian (Middle to Late Permian) brachiopod fauna from the Juripu formation in the Yarlung-Zangbo suture zone, Southern Tibet, China

A brachiopod fauna of 16 species belonging to 11 genera and three genera and species indeterminate from the middle and upper parts of the Juripu Formation in the Yarlung-Zangbo (Indus-Tsangbo) Suture zone (=Yarlung-Zangbo River zone), southern Tibet, is described and figured for the first time. A new species, Taeniothaerus zhongbaensis, is described. The fauna is comparable with that in the Kalabagh Member of the Wargal Formation of the Salt Range, Pakistan, and is considered to be most likely Capitanian (late Guadalupian/Middle Permian) to Wuchiapingian (early Lopingian/early Late Permian) in age, as indicated by the majority of the brachiopod species and by being constrained by an underlying fusulinacean fauna (Parafusulina Zone) and an overlying ammonite fauna (Cyclolobus fauna).