Palynology and dinoflagellate counts of surface sediments, and sea surface conditions, of sites in the North Pacific

Palynological analyses were performed on 53 surface sediment samples from the North Pacific Ocean, including the Bering and Okhotsk Seas (37-64°N, 144°E-148°W), in order to document the relationships between the dinocyst distribution and sea-surface conditions (temperatures, salinities, primary productivity and sea-ice cover). Samples are characterized by concentrations ranging from 18 to 143816 cysts/cm**3 and the occurrence of 32 species. A canonical correspondence analysis (CCA) was carried out to determine the relationship between environmental variables and the distribution of dinocyst taxa. The first and second axes represent, respectively, 47% and 17.8% of the canonical variance. Axis 1 is positively correlated with all parameters except to the sea-ice and primary productivity in August, which are on the negative side. Results indicate that the composition of dinocyst assemblages is mostly controlled by temperature and that all environmental variables are correlated together. The CCA distinguishes 3 groups of dinocysts: the heterotrophic taxa, the genera Impagidinium and Spiniferites as well as the cyst of Pentapharsodinium dalei and Operculodinium centrocarpum. Five assemblage zones can be distinguished: 1) the Okhotsk Sea zone, which is associated to temperate and eutrophic conditions, seasonal upwellings and Amur River discharges. It is characterized by the dominance of O. centrocarpum, Brigantedinium spp. and Islandinium minutum; 2) the Western Subarctic Gyre zone with subpolar and mesotrophic conditions due to the Kamchatka Current and Alaska Stream inflows. Assemblages are dominated by Nematosphaeropsis labyrinthus, Pyxidinopsis reticulata and Brigantedinium spp.; 3) the Bering Sea zone, depicting a subpolar environment, influenced by seasonal upwellings and inputs from the Anadyr and Yukon Rivers. It is characterized by the dominance of I. minutum and Brigantedinium spp.; 4) the Alaska Gyre zone with temperate conditions and nutrient-enriched surface waters, which is dominated by N. labyrinthus and Brigantedinium spp. and 5) the Kuroshio Extension-North Pacific-Subarctic Current zone characterized by a subtropical and oligotrophic environment, which is dominated by O. centrocarpum, N. labyrinthus and warm taxa of the genus Impagidinium. Transfer functions were tested using the modern analog technique (MAT) on the North Pacific Ocean (= 359 sites) and the entire Northern Hemisphere databases ( = 1419 sites). Results confirm that the updated Northern Hemisphere database is suitable for further paleoenvironmental reconstructions, and the best results are obtained for temperatures with an accuracy of +/-1.7 °C.

Pollen and macroremains from profiles from site Osterholz and Elm

The biostratigraphic classification of the Pleistocene in north-western and central Europe is still insufficiently known, in spite of numerous geological and vegetation-history investigations. The question is not even clear, for example, how often a warm-period vegetation with thermophilous trees such as Quercus, Ulmus, Tilia, Carpinus etc could develop here. In past years, on the basis of several geological and vegetation-history findings, suspicion has often been expressed that some of the classical stages of the Pleistocene could include more warm periods than heretofore assumed, and as a result of recent investigations the period between the Waal and Holstein interglacials seems to include at least two warm periods, of which the Cromer is one. This paper contributes to this problem. The interglacial sediments coming from the Elm-Mountains near Brunswick and from the Osterholz near Elze - both within the limits of the German Mittelgebirge - were investigated by pollen analysis. In both cases a Pinus-Betula zone and a QM zone were found. The vegetation development of the Pinus-Betula zone is characterized in both sequences by the early appearance of Picea. Because of strong local influence at the Osterholz a detailed correlation is difficult. However, vegetation development at the time of the QM zone at both sites was similar; it is especially characterized by the facts that Ulmus clearly migrated to the site earlier than Quercus and was very abundant throughout this time. Furthermore, both diagrams show very low amounts of Corylus. The interglacial of the Osterholz shows in addition to the above; a Carpinus-QM-Picea-zone in which Eucommia reaches a relative high value and in the upper of which Azolla filiculoides was also found. The similarity of vegetation development justifies acceptance of the same age for the occurrences. A comparison of the vegetation development at the Elm and the Osterholz with those of the Eem, Holstein, Waal, and Tegelen warm periods as well as with all the Cromer sites so far investigated shows that only a correlation with the Cromer Complex is possible. This correlation is supported by the geologic relations in the Osterholz (the deposit is overlain by Elster till). Therefore the till-like material with Scandinavian rock fragments underlying the deposit at Elm is of particular interest. The 'Rhume' interglacial beds at Bilshausen, only 60 km south of Osterholz, is also assigned to the Cromer complex, but the two deposits cannot be of the same age because the vegetation development differs. Therefore the Cromer complex must include at least two warm periods. Further conclusions about the relative stratigraphic position of these two occurrences and correlations of other Cromer sites are at this time not possible, however.

Pollen and diatom analysis on varved sediments in Lake Jues (Harz Mountains, Germany)

Studies combining sedimentological and biological evidence to reconstruct Holocene climate beyond the major changes, and especially seasonality, are rare in Europe, and are nearly completely absent in Germany. The present study tries to reconstruct changes of seasonality from evidence of annual algal successions within the framework of well-established pollen zonation and 14C-AMS dates from terrestrial plants. Laminated Holocene sediments in Lake Jues (10°20.70' E, 51°39.30' N, 241 m a.s.l.), located at the SW margin of the Harz Mountains, central Germany, were studied for sediment characteristics, pollen, diatoms and coccal green algae. An age model is based on 21 calibrated AMS radiocarbon dates from terrestrial plants. The sedimentary record covers the entire Holocene period. Trophic status and circulation/stagnation patterns of the lake were inferred from algal assemblages, the subannual structure of varves and the physico-chemical properties of the sediment. During the Holocene, mixing conditions alternated between di-, oligo- and meromictic depending on length and variability of spring and fall periods, and the stability of winter and summer weather. The trophic state was controlled by nutrient input, circulation patterns and the temperature-dependent rates of organic production and mineralization. Climate shifts, mainly in phase with those recorded from other European regions, are inferred from changing limnological conditions and terrestrial vegetation. Significant changes occurred at 11,600 cal. yr. BP (Preboreal warming), between 10,600 and 10,100 cal. yr. BP (Boreal cooling), and between 8,400 and 4,550 cal. yr. BP (warm and dry interval of the Atlantic). Since 4,550 cal. yr. BP the climate became gradually cooler, wetter and more oceanic. This trend was interrupted by warmer and dryer phases between 3,440 and 2,850 cal. yr. BP and, likely, between 2,500 and 2,250 cal. yr. BP.

Flux data in the Greenland Basin

Pelagic processes and their relation to vertical flux have been studied in the Norwegian and Greenland Seas since 1986. Results of long-term sediment trap deployments and adjoining process studies are presented, and the underlying methodological and conceptional background is discussed. Recent extension of these investigations at the Barents Sea continental slope are also presented. With similar conditions of input irradiation and nutrient conditions, the Norwegian and Greenland Seas exhibit comparable mean annual rates of new and total production. Major differences can be found between these regions, however, in the hydrographic conditions constraining primary production and in the composition and seasonal development of the plankton. This is reflected in differences in the temporal patterns of vertical particle flux in relation to new production in the euphotic zone, the composition of particles exported and in different processes leading to their modification in the mid-water layers. In the Norwegian Sea heavy grazing pressure during early spring retards the accumulation of phytoplankton stocks and thus a mass sedimentation of diatoms that is often associated with spring blooms. This, in conjunction with the further seasonal development of zooplankton populations, serves to delay the annual peak in sedimentation to summer or autumn. Carbonate sedimentation in the Norwegian Sea, however, is significantly higher than in the Greenland Sea, where physical factors exert a greater control on phytoplankton development and the sedimentation of opal is of greater importance. In addition to these comparative long-term studies a case study has been carried out at the continental slope of the Barents Sea, where an emphasis was laid on the influence of resuspension and across-slope lateral transport with an analysis of suspended and sedimented material.

Pollen profile and age determination of sediment core Schwanengrabenrinne, Brandenburg, Germany

In der Döberitzer Heide nördlich von Potsdam wurden vegetationsgeschichtliche Untersuchungen durchgeführt. Das Untersuchungsgebiet befindet sich im östlichen Teil der Nauener Platte, die bisher vegetationsgeschichtlich weitgehend unerforscht war. In sechs verschiedenen Mooren wurden acht Bohrungen niedergebracht. Die Bohrkerne wurden stratigraphisch und pollenanalytisch untersucht und für die Radiocarbondatierung beprobt. Die Pollendiagramme ermöglichen die Rekonstruktion der Vegetationsentwicklung der terrestrischen Standorte und der Moore in der Döberitzer Heide in den letzten 14.000 Jahren. Neben einer Revision der Gliederungsprinzipien der spätglazialen Vegetationsentwicklung Brandenburgs und einer vergleichenden Betrachtung der Moorentwicklung in der Döberitzer Heide wurde besonderes Augenmerk auf die Geschichte des Döberitzer Lindenwaldes gerichtet, der einen Sonderfall in der brandenburgischen Vegetation darstellt. Die Untersuchungen boten die Möglichkeit, die Ursachen seiner Entstehung zu klären, Aussagen zu den Perspektiven seiner Entwicklung zu treffen und mögliche Entwicklungspotentiale von Lindenwäldern im Land Brandenburg aufzuzeigen.

Pollen profiles from three former lakes in the Lake Garda area

1. Late glacial and postglacial sediments from three former lakes in the Lake Garda area (Southern Alps) were investigated. 2. The pollen diagram from Bondone (1550 m) shows an older phase rich in NAP. A younger one corresponds with the Younger Dryas time according to two radiocarbon determinations. In the Preboreal no climatic deterioration could be found. 3. At first plants, which are nowadays typical for snow-ground, pioneer and dwarf shrub associations, immigrated into the surroundings of Bondone. In Alleröd times larch and pine appeared as the first trees. At the beginning of the Preboreal dense forest existed in that region. During the Alleröd timber line was at about 1500 m. 4. In the pollen diagrams from Saltarino (194 m) and Fiavè (654 m) an oldest period rich in NAP is followed by two stadial and two interstadial phases. Tree birches and larches immigrated during the oldest interstadial phase. 5. In the case of Saltarino and Fiavè only a preliminary dating could be made. A correlation seems to be possible with diagrams published by Zoller as well as with the diagram of Bondone. Discrepances in dating, which arise then, are discussed. According to the two possibilities of dating the youngest stadial is synchronous either with the so-called Piottino stadial or the Younger Dryas time. Consequently the oldest interstadial phase of Saltarino corresponds either with the Bölling or with a pre-Bölling interstadial. The last possibility seems to be more probable. 6. In the southern part of the Lake Garda area reforestation was preceded by a long shrub phase mainly with Juniperus. At about 650 m there was a period with Pinus mugo and only with a small amount of Juniperus before reforestation. A phase with Betula nana well known from areas north of the Alps could nowhere be found. 7. In the area under study larch appeared as the first tree. Lateron it has been the most important constituent of the forests near timber line. Birch, which plays an important role as a pioneer tree in Denmark - for instance at the transition of the pollen zones III/IV - as well as in Southern Germany during Bölling time, was of less importance at the southern border of the Alps. In that area the spreading of Pinus occurred very early causing dense forests. 8. During the last stadial phase (probably Younger Dryas time) dense forests with Pinus and Larix existed at 650 m. In the lower part of the Lake Garda area, however, both thermophilous trees as Quercus and herbs frequently occurred. This leads to the conclusion that during this time tree growth was limited by dryness in lower altitudes of the border of the Southern Alps. Pinus and Juniperus, however, do not show higher values in this period, a fact which cannot yet be explained. 9. A list of plants, which were found in the sediments, is compiled. Helodium lanatum, Dictamnus albus, Mercurialis cf. ovata, Buxus, Cerinthe cf. minor, Onosma, Anthericum and Asphodelus albus are findings, which are of special interest for the history of the flora of that region.

Culture experiments to evaluate CO2 effects on growth and morphology of the coccolithophore Pleurochrysis cf. pseudoroscoffensis

In the framework of the Italian project CO2 Monitor, two culture experiments were carried out in vertical closed photobioreactors with Pleurochrysis cf. pseudoroscoffensis Gayral & Fresnel 1983, a coccolithophore isolated from the Gulf of Trieste (North Adriatic Sea). The aim of this study was to investigate the effects induced by pH variations due to CO2 emissions on its growth and morphology. Two experiments were carried out with two different CO2 concentrations (1 and 2%). Growth and cell size in light microscopy, morphology and coccolith size in scanning electron microscopy, particulate nitrogen (PN) and particulate inorganic and organic carbon (PIC and POC) content of the coccolithophore were investigated during the light and dark phases. Dissolved inorganic nutrient (nitrate and phosphate) concentrations and pH of the medium and the presence of heterotrophic prokaryotes (HP) were monitored as well.