Diversification in the South American Pampas: the genetic and morphological variation of the widespread Petunia axillaris complex (Solanaceae)

Understanding the spatiotemporal distribution of genetic variation and the ways in which this distribution is connected to the ecological context of natural populations is fundamental for understanding the nature and mode of intraspecific and, ultimately, interspecific differentiation. The Petunia axillaris complex is endemic to the grasslands of southern South America and includes three subspecies: P.a.axillaris, P.a.parodii and P.a.subandina. These subspecies are traditionally delimited based on both geography and floral morphology, although the latter is highly variable. Here, we determined the patterns of genetic (nuclear and cpDNA), morphological and ecological (bioclimatic) variation of a large number of P.axillaris populations and found that they are mostly coincident with subspecies delimitation. The nuclear data suggest that the subspecies are likely independent evolutionary units, and their morphological differences may be associated with local adaptations to diverse climatic and/or edaphic conditions and population isolation. The demographic dynamics over time estimated by skyline plot analyses showed different patterns for each subspecies in the last 100000years, which is compatible with a divergence time between 35000 and 107000years ago between P.a.axillaris and P.a.parodii, as estimated with the IMa program. Coalescent simulation tests using Approximate Bayesian Computation do not support previous suggestions of extensive gene flow between P.a.axillaris and P.a.parodii in their contact zone.

Variations in Ice Rafted Detritus on Beaches in the South Shetland Islands: A Possible Climate Proxy

Raised beach ridges on Livingston Island of the South Shetland Islands display variations in both quantity and source of ice rafted detritus (IRD) received over time. Whereas the modem beach exhibits little IRD, all of which is of local origin, the next highest beach (similar to250 C-14 yr BP) has large amounts, some of which comes from as far away as the Antarctic Peninsula. Significant quantities of IRD also were deposited similar to 1750 C-14 yr BP. Both time periods coincide with generally cooler regional conditions and, at least in the case of the similar to250 yr old beach, local glacial advance. We suggest that the increases in ice rafting may reflect periods of greater glacial activity, altered ocean circulation, and/or greater iceberg preservation during the late Holocene. Limited IRD and lack of far-travelled erratics on the modem beach are both consistent with the ongoing warming trend in the Antarctic Peninsula region.

Reduction in Northward Incursions of the South Asian Monsoon Since ~1400 AD Inferred from a Mt. Everest Ice Core

A highly resolved Mt. Everest ice core reveals a decrease in marine and increase in continental air masses related to relatively high summer surface pressure over Mongolia, and reduction in northward incursions of the summer South Asian monsoon since similar to 1400 AD. Previously published proxy records from lower sites south of the Himalayas indicate strengthening of the monsoon since this time. These regional differences are consistent with a south north seesaw in convective activity in the Asian monsoon region, and reflect a southward shift in the mean summer position of the monsoon trough since similar to 1400 AD. The change in monsoonal circulation at 1400 AD is synchronous with a reduction in solar irradiance and the onset of the LIA. This demonstrates a hemispheric scale circulation reorganization at this time, and the potential for future large shifts in monsoonal circulation.

A Census of Eddy Activities in the South China Sea During 1993-2007

Numerous mesoscale eddies occur each year in the South China Sea (SCS), but their statistical characteristics are still not well documented. A Pacific basin-wide three dimensional physical-biogeochemical model has been developed and the result in the SCS subdomain is used to quantify the eddy activities during the period of 1993-2007. The modeled results are compared with a merged and gridded satellite product of sea level anomaly by using the same eddy identification and tracking method. On average, there are about 32.9 +/- 2.4 eddies predicted by the model and 32.8 +/- 3.4 eddies observed by satellite each year, and about 52% of them are cyclonic eddies. The radius of these eddies ranges from about 46.5 to 223.5 km, with a mean value of 87.4 km. More than 70% of the eddies have a radius smaller than 100 km. The mean area covered by these eddies each year is around 160,170 km(2), equivalent to 9.8% of the SCS area with water depths greater than 1000 m. Linear relationships are found between eddy lifetime and eddy magnitude and between eddy vertical extent and eddy magnitude, showing that strong eddies usually last longer and penetrate deeper than weak ones. Interannual variations in eddy numbers and the total eddy-occupied area indicate that eddy activities in the SCS do not directly correspond to the El Nino-Southern Oscillation events. The wind stress curls are thought to be an important but not the only mechanism of eddy genesis in the SCS.

Kuroshio Intrusion and the Circulation in the South China Sea

The Princeton Ocean Model is used to study the circulation in the South China Sea (SCS) and its seasonal transition. Kuroshio enters ( leaves) the SCS through the southern ( northern) portion of the Luzon Strait. The annually averaged net volume flux through the Luzon Strait is similar to2 Sv into the SCS with seasonal reversals. The inflow season is from May to January with the maximum intrusion of Kuroshio water reaching the western SCS during fall in compensation of summertime surface offshore transport associated with coastal upwelling. From February to April the net transport reverses from the SCS to the Pacific. The intruded Kuroshio often forms an anticyclonic current loop west of the Luzon Strait. The current loop separates near the Dongsha Islands with the northward branch continuously feeding the South China Sea Warm Current (SCSWC) near the shelf break and the westward branch becoming the South China Sea Branch of Kuroshio on the slope, which is most apparent in the fall. The SCSWC appears from December to February on the seaward side of the shelf break, flowing eastward against the prevailing wind. Diagnosis shows that the onshore Ekman transport due to northeasterly monsoon generates upwelling when moving upslope, and the particular distributions of the density and sea level associated with the cross shelf motion supports the SCSWC.

A 13,600-year diatom oxygen isotope record from the South Carpathians (Romania): Reflection of winter conditions and possible links with North Atlantic circulation changes

This study provides a continuous lateglacial and Holocene record of diatom silica oxygen isotope changes (delta O-18(DIAT)) in a subalpine lake sediment sequence obtained from the Retezat Mts (Taul dintre Brazi, 1740 m a.s.l.). This through-flow, shallow, high-altitude lake with a surface area of only 0.4 ha has short water residence time and is predominantly fed by snowmelt and rainwater. Its delta O-18(DIAT) record principally reflects the oxygen isotope composition of the winter and spring precipitation, as diatom blooms occur mainly in the spring and early summer. Hence, changes in delta O-18(DIAT) are interpreted as seasonal scale changes: in the amount of winter precipitation. Low oxygen isotope values (27-28.5 parts per thousand) occurred during the lateglacial until 12,300 cal BP, followed by a sharp increase thereafter. In the Holocene delta O-18(DIAT) values ranged from 29 to 31 parts per thousand until 3200 cal BP, followed by generally lower values during the late Holocene (27-30 parts per thousand). Short-term decreases in the isotopic values were found between 10,140-9570, 9000-8500, 7800-7300, 6300-5800, 5500-5000 and at 8015, 4400, 4000 cal BP. After 3200 cal BP a decreasing trend was visible with the lowest values between 3100-2500 and after 2100 cal BP The general trend in the record suggests that contribution of winter precipitation was generally lower between 11,680 and 3200 cal BP, followed by increased contribution during the last millennia. The late Holocene decrease in delta O-18(DIAT) shows good agreement with the speleothem delta O-18, lake level and testate amoebae records from the Carpathian Mountains that also display gradual delta O-18 decrease and lake level/mire water table level rise after 3200 cal BR Strong positive correlation with North Atlantic circulation and solar activity proxies, such as the Austrian and Hungarian speleothem records, furthermore suggested that short-term increases in the isotopic ratios in the early and mid Holocene are likely connectable to high solar activity phases and high frequency of positive North Atlantic Oscillation indexes that may have resulted in decreased winter precipitation in this region.

Geomorphology and evolution of the late Pleistocene to Holocene fluvial system in the south-eastern Llanos de Moxos, Bolivian Amazon

In the Bolivian Amazon several paleochannel generations are preserved. Their wide spectrum of morphologies clearly provides crucial information on the type and magnitude of geomorphic and hydrological changes within the drainage network of the Andean foreland. Therefore, in this study we mapped geomorphological characteristics of paleochannels, and applied radiocarbon and optically stimulated luminescence dating. Seven paleochannel generations are identified. Significant changes in sinuosity, channel widths and river pattern are observed for the successive paleochannel generations. Our results clearly reflect at least three different geomorphic and hydrological periods in the evolution of the fluvial system since the late Pleistocene. Changes in discharge and sediment load may be controlled by combinations of two interrelated mechanisms: (i) spatial changes and re-organizations of the drainage network in the upper catchment, and/or (ii) climate changes with their associated local to catchment-scale modifications in vegetation cover, and changes in discharge, inundation frequencies and magnitudes, which have likely affected the evolution of the fluvial system in the Llanos de Moxos. In summary, our study has revealed the enormous potential which geomorphic mapping and analysis combined with luminescence based chronologies hold for the reconstruction of the late Pleistocene to recent fluvial system in a large portion of Amazonia.

“Unconformity-Bounded” Stratigraphic Units in the South Polar Layered Deposits (Promethei Lingula, Mars)

We conducted a stratigraphic analysis of the South Polar Layered Deposits (SPLDs) in Promethei Lingula (PL, Mars) based on the identification of regional unconformities at visible and radar wavelengths. According to the terrestrial classification, this approach constrains the stratigraphy of the region and remedies the ambiguous interpretation of stratigraphy through marker layers, bypassing the problem related to the morphologic and radiometric appearance of the layers. Thus, the approach does not exclude diverse classifications, but complements them, whereas other discriminant elements are doubtful or difficult/impossible to be defined. Using this approach, we defined two stratigraphic units (or synthems: PL1 and PL2) in PL, which are morphologically different and divided by a regional unconformity (AuR1). This stratigraphic architecture implies that the geological history of PL has been conditioned by periodic changes in climate, which in turn are related to orbital variations of Mars.