Contributions à la Paléontologie du Miocène moyen continental du Bassin du Tage. III - Mammifères - Póvoa de Santarém, Pêro Filho et Chões (Secorio). Conclusions générales

This study on middle Miocene mammalian faunas from Tagus'basin deals particularly with some small mammals hitherto undescribed from Portugal, including a new Glirid (Paraglirulus scalabicensis nov. sp.); it allows an accurate datation by biostratigraphical standards, Megacricetodon crusafonti, Fahlbuschia darocensis, Cricetodon jotae being characteristic of mammalian MN6 unit, thus their age is nearly that of Sansan and Manchones (however the presence of Peridyromys hamadryas and Lagopsis verus do suggest, amidst this biozone, a somewhat later age than Sansan's); it contributes with indirect correlation data with marine formations, as underlying oyster-bearing beds most probably are in correspondance to the apogee of the same transgression that deposed near Lisbon ”schlier" facies from VI-a division (Serravalian, Blow's zones 10-13, Globorotalia meyeri zone); the diversity of mammalian assemblages is surely related to an environement with varied biotopes, whose characterisation becomes easier if account is taken of the preceding papers on mollusks (G. Truc) and Cyprinid fishes (J. Gaudant), and also according to some unpublished paleobotanical data (J. Pais). A table with a synthesis of all paleontological data so far known is presented.

Faciès, biostratigraphie et paléogéographie du Jurassique portugais

The facies distribution along the Jurassic stages in an already well established stratigraphic frame is defined for the three portuguese basins: North of Tagus, Santiago de Cacém and Algarve. The deposits are organized in two sedimentary cycles. The first one from the Liassic to Calovian shows, in the Tagus Basin, a transgression from NW which did not surpass the Meseta present limits. The iniatilly brackish deposits only changed to marine by the end of Lotharingian. The sedimentation, mainly marly during the Liassic became more calcareous since the Aalenian. During the Dogger the basin differentiated into platform deposits towards East and South and open sea zone towards West. This zone underwent a progressive reduction and, during the Callovian, two small basins were individualized: Cabo Mondego basin in the North and Serra de El-Rei-Montejunto in the South. It is from the latter that the second sedimentary cycle (Middle Oxfordian-Portlandian) developed with open sea deposits along the Sintra–Torres Vedras axis surrounded by platform and litoral brackish formations. During the first sedimentary cycle only litoral platform deposits are known in Santiago de Cacém and Algarve basins. During the second sedimentary cycle temporary sea open deposits are known in Santiago de Cacém and Central Algarve.

Les Hipparion du Portugal

This paper is concerned with Hipparion from Ribatejo, Portugal, and with the stratigraphy of the Neogene series of this region. The first two chapters are an introduction and an historical review. Paleontological study includes both a revision of the specimens accounted by ROMAN (1907) and the description of new material. Two forms were recognized, an early H. cf. primigenium, lower Vallesian in age, NM 9 mammal unit (from Archino, Vila Nova da Rainha, Aveiras de Cima), and a more advanced H. primigenium cf. melendezi. Upper Vallesian, NM 10 (possibly lowermost Turolian, NM 11) (at Azambujeira and Marmeleira). A synthesis of Middle and Upper Miocene from Ribatejo is also presented. Levels with H. p. cf. melendezi are somewhat older than «Upper Pontian», as it was previously acknowledged, they attain at the best the lowermost Turolian (approximately corresponding to «Upper Pontian»). Even higher levels may be Turolian in age, though they are not yet accurately dated. Almost all the localities are shown (tableau 11) according to its stratigraphical position; age, correspondance to mammal units from NM 5 to NM 10 (and may be also from NM 11 to NM 12), and correlation with marine formations near Lisbon are also taken in account. The stratigraphical position of localities such as Póvoa de Santarém, Quinta do Marmelal, Pero Filho, Azambujeira (lower levels), and Fonte do Pinheiro was revised; the stratigraphical position of Marmeleira was ascertained. The localities so far known correspond to NM 5 (?), NM 6, NM 8, NM 9, NM 10 and possibly to NM II and NM 12. A new interpretation (M. T. ANTUNES) of localities with oysters from Ribatejo allows a better correlation with vertebrate localities. Relationships with Serravallian transgression seem well established. Only two localities, Vila Nova da Rainha and Foz do Alviela, may possibly be correlated to V-b division of Lisbon (Langhian) with «Hispanotherium fauna». All the other localities are younger than Serravallian oyster beds. Undirect correlation shows that NM 6 localities are somewhat younger than the apogee ef Serravallian transgression (corresponding approximately to Blow's N 11 to N 13 zones based on planctonic foraminifera).

The Neogene of Portugal

After a briefhistorical introduction, this paper deals with the main concerned geotectonic units: the Lower Tagus and Alvalade basins, the Western and Southern borders, and their infillings. Most of the Neogene events and record concern areas South of the Iberian Central Chain, a nearly inverse situation as that of Paleogene times. In the most important of these units, the Lower Tagus basin, there are quite thick detrital series, mostly marine in its distal part near Lisboa (albeit with several continental intercalations), and mainly continental in its inner part. Sedimentological record is almost complete since Lowermost to Upper Miocene. The richness ofdata (paleontology, isotope chronology, paleoclimate, etc.) it gives and the possibility of direct marine-continental correlations render this basin one of the more interesting ones in Western Europe. Alvalade basin is separated from the previous one by a barrier ofPaleozoic rocks. Two transgressions events (Upper Tortonian and Messinian in age) are recorded. Active sedimentation may be correlated to Late Miocene tectonics events. In Algarve, chiefly marine units from Lower to Upper Miocene are well developped. The Lower unit (Lagos-Portimao Formation) is best exposed in Western Algarve, but desappears eastwards. Middle Miocene is not as well known, whereas Upper Miocene main outcrops are in Eastern Algarve. Cacela Formation is remarquable for its beautiful fossils. Sedimentation as a whole refletcts the tectonic activity and in special the evolution of the Algarve flexures. There is scant evidence of post-Lower Miocene volcanism, the latest known in Portugal. Pliocene has not been recognized there beyond doubt. . Miocene sediments are much less important to the North of the Central Iberian Chain. Continental beds near Leiria that yielded the well-known "Hisp anotherium fauna" are lower Middle Miocene. Pliocene corresponds to dramatic changes in paleogeography. At Setiibal Peninsula there is some evidence of a minor Lower Pliocene transgression. Continental detrital sediments, often coarse, occupy rather large areas. In Western Portugal between the Seta hal Peninsula and Pombal there is good evidence of a marine Upper Pliocene transgression, followed up by dune sands overlain by marsh clays, diatomites, lignites and boghead levels that can be partly Pleistocene in age.

Miocene lithological, foraminiferal and palynological data from the Belverde borehole (Portugal)

XVIII Jornadas de Paleontología, 2002

The Neogene of Portugal

After a brief historical introduction, this paper deals with the main concerned geotectonic units: the Lower Tagus and Alvalade basins, the Western and Southern borders, and their infillings. Most of the Neogene events and record concern areas South of the Iberian Central Chain, a nearly inverse situation as that of Paleogene times. In the most important of these units, the Lower Tagus basin, there are quite thick detrital series, mostly marine in its distal part near Lisboa (albeit with several continental intercalations), and mainly continental in its inner part. Sedimentological record is almost complete since Lowermost to Upper Miocene. The richness ofdata (paleontology, isotope chronology, paleoclimate, etc.) it gives and the possibility of direct marine-continental correlations render this basin one of the more interesting ones in Western Europe. Alvalade basin is separated from the previous one by a barrier of Paleozoic rocks. Two transgressions events (Upper Tortonian and Messinian in age) are recorded. Active sedimentation may be correlated to Late Miocene tectonics events. In Algarve, chiefly marine units from Lower to Upper Miocene are well developped. The Lower unit (Lagos-Portimão Formation) is best exposed in Western Algarve, but desappears eastwards. Middle Miocene is not as well known, whereas Upper Miocene main outcrops are in Eastern Algarve. Cacela Formation is remarquable for its beautiful fossils. Sedimentation as a whole refletcts the tectonic activity and in special the evolution of the Algarve flexures. There is scant evidence of post-Lower Miocene volcanism, the latest known in Portugal. Pliocene has not been recognized there beyond doubt. Miocene sediments are much less important to the North of the Central Iberian Chain. Continental beds near Leiria that yielded the well-known "Hisp anotherium fauna" are lower Middle Miocene. Pliocene corresponds to dramatic changes in paleogeography. At Setiibal Peninsula there is some evidence of a minor Lower Pliocene transgression. Continental detrital sediments, often coarse, occupy rather large areas. In Western Portugal between the Setúbal Peninsula and Pombal there is good evidence of a marine Upper Pliocene transgression, followed up by dune sands overlain by marsh clays, diatomites, lignites and boghead levels that can be partly Pleistocene in age.

Paleoclimatic correlations between Western european continental and marine areas Contribution of big villafranchian fauna

It is very difficult to make paleoclimatic correlations between continental and marine areas, but it is possible with biostratigraphic data. Reliable correlations can be made only between broad periods: between 3.5 and 3 Ma, around 2.4 Ma, until 1.6 Ma and after 1.6 Ma. The arid Mediterranean phases led to the disappearance of the European Villafranchian fauna (1.0 Ma).

The marine Neogene of Eastern Venezuela : a preliminary report

Proceedings of tile 1" R.C.A.N.S. Congress, Lisboa, October 1992

The Cristo Rei section (Lower Miocene). Distal fluviatile environments in a marine series, plants, vertebrates and other evidence, age

The section at Cristo Rei shows sandy beds with intercalated clayey lenses (IVb division from the Lisbon Miocene series) that correspond to a major regression event dated from between ca. 17.6 and 17 Ma. They also correspond to a distal position (relatively to the typical fluviatile facies in Lisbon), nearer the basin's axis. Geologic data and paleontological analysis (plant fossils, fishes, crocodilians, land mammals) allow the reconstruction of environments that were represented in the concerned area: estuary with channels and ox-bows; upstream, areas occupied by brackish waters where Gryphaea griphoides banks developped; still farther upstream, freshwaters sided by humid forests and low mountain subtropical forests under warm temperate and rainy conditions, as well as not far away, seasonally dry environments (low density tree or shrub cover, or steppe).

The three main marine depositional cycles of the Neogene of Portugal

The present work follows a stratigraphic model for the marine Neogene of Portugal based on the definition of three main marine sedimentary cycles. Conceptually the I, II and III Neogene Cycles can be defined as 2nd order sedimentary sequences with duration ranging from 5 to 8 Ma. The I Neogene Cycle is fully represented only in the Lower Tagus Basin. Ranging from the Early Aquitanian to the Late Burdigalian the I Neogene Cycle testify a transgressive episode in the region of Lisbon and Setúbal Peninsula. Rapid lateral facies variations suggest a shallowmarine basin. This cycle ends with an important Late Burdigalian tectonic compressive event expressed by uplift of the surrounding areas and deformation affecting the Early Miocene deposits of the Arrábida Chain. The II Neogene Cycle includes thick sedimentary sequences covering Paleozoic and Mesozoic formations in the Algarve and Alvalade-Melides regions and it extends as far north as Santarém in the Lower Tagus Basin. Mainly controlled by global eustasy, it was generated by the important positive eustatic trend that characterized the Middle Miocene worldwide to which the Portuguese continental margin acted more or less passively. This cycle ended with a second and the most important compression event starting after the end of the Serravallian affecting the entire Portuguese onshore and shelf areas. This led to an important depositional hiatus of marine sediments for more than 2.5 Ma. During the Early and the Middle Tortonian occurred the clockwise rotation of the Guadalquivir Basin. The thickmarine units deposited afterwards in this basin produced a litostatic load, which seems to have induced subsidence farther west resuming the Neogene marine sedimentation in the Cacela region (Eastern Algarve), during the Late Tortonian. This marks the beginning of the III Neogene Cycle. To the north, in the Sado Basin (Alvalade-Melides region), a similar depositional sequence starts its sedimentation during the Messinian. Further north, in the Pombal-Caldas da Rainha region, marine sedimentation started during the Late Pliocene (Piacenzian). The migration in time, from south to north for the beginning of the marine sedimentation of this cycle is interpreted as reflecting a visco-elastic propagation of the deformation from the Betic chain northwards.

Late Quaternary warm marine mollusks from Santa Maria (Azores); paleoecologic and paleobiogeographic considerations

The sub-fossil fauna from the Late Quaternary marine deposits of Santa Maria is made of more than 50 species of gastropods and bivalves, 19 of them collected recently and for the first time in the northern coast of the island (Lagoinhas Bay). The sub-fossil shells are found in deposits of beach sands, situated 2-3 meters above the present low tide. The carbonated sands from the basal part of the succession yield an autochthonous association of borers dominated by the bivalve Myoforceps aristata (Dillwin, 1817). Upwards, the marine sands contain concentrations of beach drift shells, including well-preserved supratidal and intertidal gastropods, among them a large number of Rissoidae. The bivalve fauna is dominated by disarticulated valves of Ervilia castanea (Montagu, 1803), a small infaunal coloniser of mobile sandy substrates. The composition of the fauna is made essentially of West European species, many of them common to the West Coast of Portugal. However, a few "warm guests" with West African or Caribbean affinities were also found, suggesting a close relation with some of the "Tyrrhenian" warm associations found in the Western Mediterranean.

Miocene marine mammals from Portugal. Paleogeographical and paleoecological significance

Neogene marine mammals are still incompletely known in Portugal. However, a general overview of the geographic and stratigraphic distribution of marine mammal localities in the Miocene of Portugal is already possible. An attempt of correlation between the trends shown by these distributions and the horizontal and vertical environmental shifts is presented. In general, sirenians occur in deposits representing shallow, warm, low energy aquatic environments; while cetaceans are more frequent in more open, deep and temperate marine environments.

Miocene marine-continental correlations in the Lisbon area and some discussion related to personnal experience

Sandpit exploitation near Lisbon allowed collecting of many Miocene, non marine fossils. These sands are part of the mostly marine Miocene series in the Lower Tagus basin. The particularly favourable situation led several researchers to deal with marine-continental correlations. Difficulties often concern methodologic aspects. Some poorly based interpretations exerced a lasting influence. A critical approach is presented. Analysis requires data. Methods based upon models often lead to the temptation of fitting data in order to confirm a priori conclusions, or of mixing up data as if of equal statistic value while they have not at all the same weight. Erroneous interpretations' uncritical repetition for many years "upgraded" them into absolute truth. Another point is endemism vs. europeism. Miocene mammals from Lisbon compared well with corresponding French, contemporaneous taxa, while this was apparently not true for Spanish ones. Too much accent had been put on the endemic character of Spanish, or even regional, mammalian faunas. Nationalist bias and sensationalism also weigh, albeit negatively. Meanwhile nearly all the more evident examples as the rhinoceros Hispanotherium are discredited as Iberian endemisms. Taxa may appear as endemic just because they have not yet been found elsewhere. At least for the medium to large-sized mammals, with their huge geographic distribution, faunal differences depend much more on ecology, climate and environmental conditions. Emphasis on differences may also result from researchers that are often in a precarious situation and need very much to achieve short-term, preferably sensational results. Overvalued differences may mask real similarities. Unethic and not scientific behaviour are further enhanced by "nomina nuda" tricks that may simply be a way to circunvent or cheat the Priority Rule. On the other hand, access to communication networks may present as sensational novelties items that are not new at all, misleading the audience. A new class of "science people" arose, created by the media and not by the value of their real achievements. Discussion is presented on sedimentation processes and discontinuities that are often regarded as absolute precision dating tools, as well as on some geochemical and paleomagnetic interpretations. A very good chronologie frame has been obtained for the basin under study on the basis of an impressive set of data, providing a rather detailed and accurate frame for Miocene marine-continental correlations.

Picocyanobacteria From a Clade Of Marine Cyanobium Revealed Bioactive Potential Against Microalgae, Bacteria, And Marine

The production of bioactive compounds either toxic or with pharmacological applications by cyanobacteria is well established. However, picoplanktonic forms within this group of organisms have rarely been studied in this context. In this study, the toxicological potential of picocyanobacteria from a clade of marine Cyanobium strains isolated from the Portuguese coast was examined using different biological models. First, strains were identified by applying morphological and molecular approaches and cultured under lab conditions. A crude extract and three fractions reflecting a preliminary segregation of lipophilic metabolites were tested for toxicity with the marine microalga Nannochloropsis sp., the bacteria Pseudomonas sp., the brine shrimp Artemia salina, and fertilized eggs of the sea urchin Paracentrotus lividus. No significant apparent adverse effects were noted against Artemia salina. However, significant adverse effects were found in all other assays, with an inhibition of Nannochloropsis sp. and Pseudomonas sp. growth and marked reduction in Paracentrotus lividus larvae length. The results obtained indicated that Cyanobium genus may serve as a potential source of interesting bioactive compounds and emphasize the importance of also studying smaller picoplanktonic fractions of marine cyanobacteria.