Late Cretaceous planktonic foraminiferal assemblages from the Exmouth Plateau, ODP Sites 122-762 and 122-763, eastern Indian Ocean

The evolution of planktonic foraminifera during the Late Cretaceous is marked in the Santonian by the disappearance of complex morphotypes (the marginotruncanids), and the contemporary increasing importance and diversification of another group of complex taxa, the globotruncanids. Upper Turonian to lower Campanian planktonic foraminiferal assemblages from Holes 762C and 763B (Ocean Drilling Program, Leg 122, Exmouth Plateau, 47°S palaeolatitude) were studied in detail to evaluate the compositional variations at the genus and species level based on the assumption that, in the Cretaceous oceans as in the modern, any faunal change was associated with changes in the characteristics and the degree of stability of the oceanic surface waters. Three major groups were recognised based on gross morphology, and following the assumption that Cretaceous planktonic foraminifera, although extinct, had life-history strategies comparable to those of modern planktonics: 1 - r-selected opportunists; 2 - k-selected specialists; 3 - r/k intermediate morphotypes which include all genera that display a range of trophic strategies in-between opportunist and specialist taxa. Although planktonic foraminiferal assemblages are characterised by a progressive appearance of complex taxa, this trend is discontinuous. Variation in number of species and specimens within genera has allowed recognition of five discrete intervals each of them reflecting different oceanic conditions based on fluctuations in diversity and abundance of the major morphotypes. Planktonic forms show cyclical fluctuations in diversity and abundance of cold (r-strategists) and warm taxa (k-strategists), perhaps representing alternating phases of unstable conditions (suggesting a weakly stratified upper water column in a mesotrophic environment), and well-stratified surface and near-surface waters (indicating a more oligotrophic environment). Interval 1, middle Turonian to early Coniacian in age, is dominated by the r/k intermediate morphotypes which alternate with r-strategists. These cyclical alternations are used to identify three additional subintervals. Interval 2, aged middle to late Coniacian, is characterised by the increasing number of species and relative abundance of k-strategists. After this maximum diversification the k-strategists show a progressive decrease reaching a minimum value in Interval 3 (early to late Santonian), which corresponds to the extinction of the genus Marginotruncana. In the Interval 4, latest Santonian in age, the k-strategists, represented mainly by the genera Globotruncana, increase again in diversity and abundance. The last Interval 5 (early Campanian) is dominated by juvenile globotruncanids and r-strategists which fluctuate in opposite phase. The positive peak (Interval 2) related to the maximum diversification of warm taxa (k-strategists) in the Coniacian seems to correspond to a warmer episode. It is followed by a marked decrease in the relative abundance of warm taxa (k-strategists crisis) with a minimum in the late Santonian (Interval 3), reflecting a decrease in temperature. Detailed analysis of faunal variations allows the Santonian faunal turnover to be ascribed to a cooling event strong enough to cause the extinction of the marginotruncanids.

Fossil record of Mollusca from Neogene sediments in Schleswig-Holstein, Germany

Über die Verbreitung, Gliederung und Ausbildung des Jungtertiärs im westlichen Schleswig-Holstein war bisher nicht viel bekannt. Am besten bearbeitet sind die glazial gestauchten Schollen von Morsum/Sylt. Eine Aufzählung erbohrter Miozänvorkommen mit nicht immer überzeugender Begründung lieferte H.-L. HECK 1935. S. THIELE (1941) hat die ihm bekannten Vorkommen hauptsächlich nach faziellen und petrographischen Gesichtspunkten bearbeitet. Er erkannte richtig die Stellung der Braunkohlensande. Die angekündigte palaeontologische Bearbeitung ist nicht erschienen. Eine allgemeine Übersicht über die Entwicklung des Jungtertiärs bringen W. WOLFE und H.-L. HECK 1949. W. HINSCH lieferte wertvolle Beiträge zur Molluskenfauna und zur Gliederung des Miozäns (1952, 1955). Über neue Vorkommen von Braunkohlen-Sanden berichtete E. DITTMER(1 956), eine erste Übersicht über neue Vorkommen der Hemmoorer Stufe gab derselbe Verfasser 1957.

Upper Turonian - lower Campanian planktonic foraminifera from Exmouth Plateau

A planktonic foraminiferal zonal scheme is presented for subdivision of the Upper Cretaceous pelagic carbonate sequence from southern mid-high latitudes. Definition of the zones is based on first and last occurrences of planktonic foraminifera from Ocean Drilling Program Holes 762C and 763B (Leg 122; Exmouth Plateau, south Indian Ocean). During the Late Cretaceous the studied holes were located close to 50°S and for the first time a complete sedimentary record for the mid-high latitudes was obtained. A detailed biostratigraphic analysis has allowed recognition of two new zones (Falsotruncana maslakovae Zone and Marginotruncana marianosi Zone) for the interval extending from the last occurrence of Helvetoglobotruncana helvetica to the first occurrence of Dicarinella asymetrica (upper Turonian - lower Santonian). From this study it is apparent that some low latitude (Globotruncana ventricosa, Hedbergella flandrini, Marginotruncana marianosi) and high latitude (Globigerinelloides impensus and Hedbergella sliteri) marker taxa display a vertical distribution at mid-high latitudes which is different from that known from low latitudes; moreover, one species (Heterohelix papula), overlooked at low latitudes, exhibits a restricted range that seems to be useful for chrono-biostratigraphic correlations: its appearance is suggested to coincide with the Coniacian/Santonian boundary. The proposed biozonation, which is integrated with calcareous nannofossil and magnetostratigraphic data available for the sections studied, is compared with both the low-latitude standard zonation and the planktonic foraminiferal zonal scheme for the circum-Antarctic region, in order to define a bio-chronostratigraphic scale that is useful for mid-high latitudes of the southern oceans.