A new genus Abyssogena from deep-water vents and seeps

A new genus Abyssogena is established for A. phaseoliformis (Métivier, Okutani & Ohta, 1986) and A. kaikoi (Okutani & Métivier, 1986), which were previously assigned to the genus Calyptogena Dall, 1891, and also for two new species, A. southwardae and A. novacula. The most characteristic features of Abyssogena are an elongate shell up to about 280 mm in length; a pallial line starting from the ventral margin of the anterior adductor scar; secondary pallial attachment scars developed dorsal to the pallial line; radially arranged hinge teeth with a reduced anterior cardinal tooth in the right valve; and presence of an inner ctenidial demibranch only. Abyssogena occurs in deep water from 2,985 to 6,400 m and is distributed in the Pacific and Atlantic Oceans at cold seeps along continental margins and hydrothermal vents at mid-oceanic ridges. Some species have a remarkably wide geographic distribution; A. southwardae is present throughout the Atlantic and A. phaseoliformis is present in Japan, Kuril-Kamchatka, as well as Aleutian Trenches. No fossils of Abyssogena are known.

Distribution of dinoflagellate cycst at ODP Site 123-765 (Table 1)

Site 765 contains a sequence of tropical, middle Miocene to Holocene dinoflagellate cysts. These diverse assemblages are characterized by abundant Polysphaeridium zoharyi and Spiniferites bulloideus. Abundances of Impagidinium spp. and Nematosphaeridium spp. reflect the shelf-to-slope origin of the assemblages. One new genus, Blysmatodinium, and two new species, Nematosphaeridium (?) wrennii sp. nov. and Blysmatodinium argoi, are described.

Calcareous nannofossil stratigraphy and datums of sediments from ODP Leg 198 sites

During Ocean Drilling Program Leg 198, Sites 1207, 1208, 1212, 1213, and 1214 were drilled on Shatsky Rise, coring Lower to mid-Cretaceous successions of nannofossil chalk, porcellanite, and chert. Although recovery was poor, these sites yielded an outstanding record of calcareous nannoplankton, providing valuable data concerning the evolutionary succession and paleobiogeography of the largest Cretaceous marine habitat. Mid-Cretaceous sections (Aptian-Cenomanian) were recovered at all sites, and Site 1213 includes an apparently complete Berriasian-Hauterivian section. Biostratigraphic dating is problematic in places because of the absence or rarity of zonal fossils of both Boreal and Tethyan affinity. The majority of nannofossil assemblages are relatively typical of this age, but there are clear differences that set them apart from coeval epicontinental assemblages: for example, Lithraphidites carniolensis is common to abundant throughout and was most likely an oceanic-adapted taxon; the cold- to temperate-water species Crucibiscutum salebrosum, Repagulum parvidentatum, and Seribiscutum primitivum are entirely absent, indicating the persistence of tropical, warm surface water temperatures; and the warm-water species Hayesites irregularis is common. Most striking, however, is the virtual absence of Nannoconus and Micrantholithus, both taxa that were conspicuous and often common components of many Tethyan and Atlantic nannofloras. These forms were almost certainly neritic adapted and usually absent in deep open-ocean settings away from guyots and platforms. Other Tethyan taxa are also absent or rare and sporadically distributed (e.g., Calcicalathina oblongata, Conusphaera spp., Tubodiscus verenae, and Lithraphidites bollii), and factors related to neritic environments presumably controlled their distribution. Site 1213 also records extended Early Cretaceous ranges for species previously thought to have become extinct during the Late Jurassic (e.g., Axopodorhabdus cylindratus, Hexapodorhabdus cuvillieri, and Biscutum dorsetensis), suggesting these species became Pacific-restricted prior to their extinction. Watznaueria britannica may also have been a species with Pacific affinities before reexpansion of its biogeography in the early Aptian. One new genus (Mattiolia) and thirteen new species (Zeugrhabdotus clarus, Zeugrhabdotus petrizzoae, Helicolithus leckiei, Rhagodiscus amplus, Rhagodiscus robustus, Rhagodiscus sageri, Rhagodiscus adinfinitus, Tubodiscus bellii, Tubodiscus frankiae, Gartnerago ponticula, Haqius peltatus, Mattiolia furva, and Kokia stellata) are described from the Shatsky Rise Lower Cretaceous section.

Late Cretaceous dinoflagellate cysts of ODP Hole 120-748C

At Ocean Drilling Program Hole 748C in the Southern Indian Ocean, a total of 171 Late Cretaceous dinoflagellate taxa were encountered in 38 productive samples from Cores 120-748C-27R through 120-748C-62R (407-740 mbsf). Four provisional dinoflagellate assemblage zones and five subzones were recognized based on the character of the dinoflagellate flora and the first/last occurrences of some key species. Isabelidinium korojonense and Nelsoniella aceras occur in Zone A together with Oligosphaeridium pulcherrimum and Trithyrodinium suspect urn. Zone B was delineated by the total range of Odontochitina cribropoda. Zone C was separated from Zone B by the presence of Satyrodinium haumuriense, and Zone D is dominated by new taxa. The dinocyst assemblages bear a strong affinity to Australian assemblages. Paleoenvironmental interpretations based mainly on dinocysts suggest that during the ?Santonian-Campanian to the Maestrichtian this portion of the Kerguelen Plateau was a shallow submerged plateau, similar to nearshore to offshore to upper slope environments with water depths of tens to hundreds of meters, but isolated from the major continents of the Southern Hemisphere. Starting perhaps in the late Cenomanian (Mohr and Gee, 1992, doi:10.2973/odp.proc.sr.120.196.1992), the Late Cretaceous transgression over the plateau reached its maximum during the late Campanian. The plateau may have been exposed above sea level and subjected to weathering during the latest Maestrichtian. The studied dinocyst assemblages characterized by species of Amphidiadema, Nelsoniella, Satyrodinium, and Xenikoon together with abundant Chatangiella (the large-size species) and Isabelidinium suggest that a South Indian Province (tentatively named the Helby suite) may have existed during the Campanian-Maestrichtian in comparison with the other four provinces of Lentin and Williams. One new genus, three new species, and two new subspecies of dinocysts are described.

Diatom abundances in ODP Leg 114 holes

The practically continuous, paleomagnetically dated late Gauss-Brunhes sediment profiles of ODP Sites 699 and 701, south of the present Polar Front Zone (PFZ), and Site 704, north of the present PFZ, are used for a high-resolution study of abundance fluctuations of eight stratigraphic marker species in space and time. Ecological restrictions and preferences of the diatom species Hemidiscus karstenii, Actinocyclus ingens f. planus, Thalassiosira elliptipora, Thalassiosira kolbei, Thalassiosira vulnifica, Simonseniella barboi, Cosmiodiscus insignis, and Nitzschia weaveri are deduced. The ages of their first abundant appearance datums (FAAD), last-appearance datums (LAD), and last abundant appearance datums (LAAD) at the three sites are determined. The interpolated datum ages agree relatively well with those determined by other authors, if one interprets most of their LADs as LAADs. FAADs and LAADs produce more accurate datums than LADs. For the late Matuyama (younger than approximately 2.0 Ma), when PFZ fluctuations effected all three site sites, the datum ages determined agree within the methodically caused limits of accuracy for each datum. For the early Matuyama (older than approximately 2.0 Ma) the results can be interpreted as either that the ages of the FAAD of T. kolbei and LAAD of T. vulnifica datums determined at Sites 699 and 701 are more reliable or that these datums are diachronous between these two sites and Site 704. Such a diachroneity could be caused by different paleoceanographic conditions (stable subantarctic conditions over Site 704 and stable antarctic conditions over Sites 699 and 701). A few taxonomic changes were necessary. One new genus is defined (Simonseniella gen. nov.) and five new combinations are proposed: Simonseniella barboi (Brun) comb, nov., Simonseniella praebarboi (Schrader) comb, nov., Simonseniella curvirostris (Jousé) comb, nov., Thalassiosira elliptipora (Donahue) comb, nov., and Thalassiosira vulnifica (Gombos) comb. nov.

Oligocene-Miocene calcareous nannofossils in ODP Leg 149 holes

During Ocean Drilling Program (ODP) Leg 149, five sites were drilled on the Iberia Abyssal Plain in the northeastern Atlantic Ocean. Both Mesozoic and Cenozoic sediments were recovered. Oligocene to Miocene sediments were cored at deepwater Sites 897, 898, 899, and 900. Except for a few intervals, occurrences of generally abundant and well-preserved calcareous nannofossils suggest that the deposition of the turbidite-type sediments occurred above the calcite compensation depth (CCD). One major unconformity in the middle late Miocene is present. Detailed quantitative analyses of calcareous nannofossils are used to determine the changes occurring among the nannoflora in relation to sea-level variation. A succession of 89 biohorizons from the early Oligocene to the late Miocene are defined by combining the biostratigraphic results of the four sites studied in the Iberia Abyssal Plain. One new genus and eight new species are described: Camuralithus, Camuralithus pelliculatus, Ericsonia detecta, Helicosphaera limasera, Sphenolithus akropodus, Sphenolithus aubryae, Sphenolithus cometa, Reticulofenestra circus, and Syracosphaera lamina. Two new variations and seven new combinations are also introduced.

Nematodes and meiofauna in sediments obtained during Belgica cruise BG06/13, north-east Atlantic

The Gollum Channels and Whittard Canyon (NE Atlantic) are two areas that receive high input of organic matter and phytodetritus from euphotic layers, but they are typified by different trophic and hydrodynamic conditions. Sediment biogeochemistry was analysed in conjunction with structure and diversity of the nematode community and differences were tested between study areas, water depths (700 m vs 1000 m), stations, and sediment layers. The Gollum Channels and Whittard Canyon harboured high meiofauna abundances (1054-1426 ind. 10 cm**-2) and high nematode diversity (total of 181 genera). Next to enhanced meiofauna abundance and nematode biomass, there were signs of high levels of organic matter deposition leading to reduced sedimentary conditions, which in turn structured the nematode community. Striking in this respect was the presence of large numbers of 'chemosynthetic' Astomonema nematodes (Astomonema southwardorum, Order Monhysterida, Family Siphonolaimidae). This genus lacks a mouth, buccal cavity and pharynx and possesses a rudimentary gut containing internal, symbiotic prokaryotes which have been recognised as sulphur-oxidising bacteria. Dominance of Astomonema may indicate the presence of reduced environments in the study areas, which is partially confirmed by the local biogeochemical environment. The nematode communities were mostly affected by sediment layer differences and concomitant trophic conditions rather than other spatial gradients related to study area, water depth or station differences, pointing to small-scale heterogeneity as the main source of variation in nematode structure and function. Furthermore, the positive relation between nematode standing stocks, and quantity and quality of the organic matter was stronger when hydrodynamic disturbance was greater. Analogically, this study also suggests that structural diversity can be positively correlated with trophic conditions and that this relation is tighter when hydrodynamic disturbance is greater.

Early Cretaceous planktonic foraminifera from the Tethys

Early Cretaceous planktonic foraminiferal assemblages include rare planispiral and pseudoplanispiral morphotypes with elongate chambers that BouDagher-Fadel et al. assigned to Schackoina or accommodated in the new genus Claviblowiella. New findings of well-preserved planktonic foraminiferal faunas from the Lesches en Diois (SE France) section, the Cismon core (NE Italy), the Calabianca (NW Sicily) section and the Upper Aptian of Deep Sea Drilling Project (DSDP) Site 545 drilled off Morocco, have allowed a better understanding of the morphological features of these rare, unevenly distributed taxa. Our data demonstrate that each small planispiral species with globular chambers has a corresponding "clavate" morphotype which (as the "normal" forms) exhibits a smooth, finely perforate wall. Consequently, the latter have been assigned here to the genus Globigerinelloides and treated as subspecies of the "non-clavate" taxa. The (clavate) subspecies belonging to the genus Globigerinelloides here retained are G. duboisi sigali Longoria, G. maridalensis elongatus subsp. nov., G. blowi lobatus subsp. nov. and G. paragottisi clavatus subsp. nov., while Globigerinelloides minai Obregòn de la Parra is not retained. In addition, a new genus, Pseudoschackoina, type species Planomalina saundersi Bolli (senior synonym of Hastigerinoides cepedai Obregòn de la Parra, has been formalised for individuals possessing elongate, pointed, laterally compressed chambers, bearing tubulospines arranged on a pseudoplanispiral (dysaxial) coiling mode. Stratigraphically, in the sections studied the first taxon to appear is Pseudoschackoina saundersi, in the uppermost part of the Selli Level (=OAE1a), immediately followed, just above the OAE1a, by all the "clavate" globigerinelloidids. Regarding the last occurrences, Pseudoschackoina saundersi and G. maridalensis elongatus disappear in the lower part of the Globigerinelloides algerianus Zone, Globigerinelloides paragottisi clavatus at the top of the same zone, while Globigerinelloides blowi lobatus and G. duboisi sigali range up to the lower part of the Ticinella bejaouaensis Zone.

Distribution of calcareous nannofossils in the Miocene interval of DSDP Hole 94-608 in the north Atlantic (Fig. 1)

DSDP North Atlantic Site 608 yielded an excellent Miocene pelagic section which affords a further opportunity for elucidating the chronology of the calcareous nannofossil succession in the framework of magnetostratigraphic control. Most of the conventional (zonal) markers have been documented for this site and some of the earlier results are confirmed and refined. In addition several unconventional and less known markers have been added. The first two are the highest (last) occurrence of Sphenolithus delphix and Sphenolithus capricornutus at 23.6 Ma, which is immediately above the Oligocene-Miocene boundary as identified by the last occurrence of Reticulofenestra bisecta at 23.7 Ma. The next unconventional datum is the highest (last) occurrence of Ilselithina fusa at 22.8 Ma, which is also the highest (last) occurrence of Helicosphaera recta. Calcidiscus tropicus' lowest (first) occurrence is at 19.5 Ma, which is also the lowest occurrence of Sphenolithus belemnos, and Calcidiscus leptoporus' lowest (first) occurrence coincides with that of Sphenolithus heteromorphus at 18.5 Ma. Sphenolithus dissimilis' highest (last) occurrence is at 18.2 Ma and the Calcidiscus premacintyrei lowest (first) and highest (last) occurrences are, respectively, at 17.7 and 11.7 Ma. Discoaster braarudii occurs from 11.6 to 11.3 Ma and its highest (last) occurrence corresponds to that of Cyclicargolithus floridanus. Minylitha convallis occurs from 9.0 to 6.9 Ma. Within the range of Minylitha, at 8.0 Ma, a major shift occurs in reticulofenestrid placoliths from dominantly large (Reticulofenestra pseudoumbilicus) and medium size (Reticulofenestra minutula) species below to significant numbers of very small species (Dictyococcites productus and Gephyrocapsa) above. This is interpreted to be a major, though perhaps seasonal, change of productivity of the North Atlantic at Site 608. A new genus and species Cryptococcolithus takayamae, is described and a variety, Reticulofenestra pseudoumbilicus var. amplus is identified.

Distribution of desmoscolecida (Nematoda) in surface sediments of the Iberian Deep-Sea, North Atlantic (Table 1)

1. Desmoscolecida from the continental slope and the deep-sea bottom (59-4354 m) off the Portuguese and Moroccan coasts are described. 18 species were identified: Desmoscolex bathyalis sp. nov., D. chaetalatus sp. nov., D. eftus sp. nov., D. galeatus sp. nov., D. lapilliferus sp. nov., D. longisetosus Timm, 1970, D. lorenzeni sp. nov., D. perspicuus sp. nov., D. pustulatus sp. nov., Quadricoma angulocephala sp. nov., Q. brevichaeta sp. nov., Q. iberica sp. nov., Q. loricatoides sp. nov., Tricoma atlantica sp. nov., T. bathycola sp. nov., T. beata sp. nov., T. incomposita sp. nov., T. meteora sp. nov., T. mauretania sp. nov. 2. The following new terms are proposed: "Desmos" (ring-shaped concretions consisting of secretion and concretion particles), "desmoscolecoid" and "tricomoid" arrangement of the somatic setae, "regelmaessige" (regular), "unregelmaessige" (irregular), "vollstaendige" (complete) and "unvollstaendige" (incomplete) arrangement of somatic seta (variations in the desmoscolecoid arrangement of the somatic setae). The length of the somatic setae is given in the setal pattern. 3. Desmoscolecida identical as to genus and species exhibit no morphological differences even if forthcoming from different bathymetrical zones (deep sea, sublitoral, litoral) or different environments (marin, freshwater, coastal subsoil water, terrestrial environment). 4. Lorenzen's (1969) contention that thearrangement of the somatic setae is more significant for the natural relationships between the different genera of Desmoscolecida than other characteristics is further confirmed. Species with tricomoid arrangement of somatic setae are regarded as primitive, species with desmoscolecoid arrangement of somatic setae are regarded as more advanced. 5. Three new genus are established: Desmogerlachia gen. nov., Desmolorenzenia gen. nov. and Desmofimmia gen. nov. - Protricoma Timm, 1970 is synonymized with Paratricoma Gerlach, 1964 and Protodesmoscolex Timm, 1970 is synonymized with Desmoscolex Claparede,1863. 6. Checklists of all species of the order Desmoscolecida and keys to species of the subfamilies Tricominae and Desmoscolecinae are provided. 7. The following nomenclatorial changes are suggested: Desmogerlachia papillifer (Gerlach, 1956) comb. nov., D .pratensis (Lorenz, 1969) comb. nov., Desmotimmia mirabilis (Timm, 1970) comb. nov., Paratricoma squamosa (Timm, 1970) comb. nov., Desmolorenzenia crassicauda (Timm, 1970) comb. nov., D. desmoscolecoides (Timm, 1970) comb. nov., D. eurycricus (Filipjev, 1922) comb. nov., D. frontalis (Gerlach, 1952) comb. nov., D. hupferi (Steiner, 1916) comb. nov., D. longicauda (Timm, 1970) comb. nov., D. parva (Timm, 1970) comb. nov., D. platycricus (Steiner, 1916) comb. nov., D. viffata (Lorenzen, 1969) comb. nov., Desmoscolex anfarcficos (Timm, 1970) comb. nov.

New Pliocene and Pleistocene acritarchs from the North Atlantic, Arctic Ocean and Bering Sea

Acritarchs have received limited attention in palynological studies of the Cenozoic, although they have much potential both for refining Neogene and Quaternary stratigraphy, especially in mid- and high northern latitudes, and developing palaeoceanographical reconstructions. Here we formally describe and document the stratigraphical and palaeotemperature ranges (from foraminiferal Mg/Ca) of four new acritarch species: Cymatiosphaera? aegirii sp. nov., Cymatiosphaera? fensomei sp. nov., Cymatiosphaera? icenorum sp. nov. and Lavradosphaera canalis sp. nov. In reviewing the stratigraphical distributions of all species of the genus Lavradosphaera De Schepper & Head, 2008, we demonstrate their correlation potential between the North Atlantic and Bering Sea in the Pliocene. Additionally, Lavradosphaera lucifer De Schepper & Head, 2008 and Lavradosphaera canalis sp. nov., while not themselves overlapping stratigraphically, have morphological intermediates that do partially overlap and may represent an evolutionary trend consequent upon climate cooling in the Late Pliocene. Finally, we show that the highest abundances of the acritarchs presented here were living in the eastern North Atlantic, in surface-water temperatures not very different from today.