Oligocene and Neogene radiolarians from the Labrador Sea

Two sites in the Labrador Sea and one site in Baffin Bay were drilled during Leg 105. Radiolarians were recovered at all three sites, although at Site 645 (Baffin Bay), radiolarians were present in useful numbers only in the mudline sample. Radiolarians of late Neogene age were recovered at Site 646 south of Greenland, while early Oligocene and early Miocene radiolarians were recovered from the Labrador Sea at Site 647. In Site 646, radiolarian and other coarse-fraction abundances vary dramatically from sample to sample and may reflect deep-water depositional processes as well as changes in surface-water conditions. Site 647 siliceous microfossils reach their peak abundance and preservation in Core 105-647A-25R and decline gradually upward into the lower Miocene (Cores 105-647A-13R and -14R). Siliceous microfossil abundances in counts of the > 38-µm Carbonate-free coarse fraction from the siliceous interval are correlated to each other, but not to the abundance of nonbiogenic coarse-fraction components. Radiolarian abundances in specimens per gram (but not diatom abundances) are correlated to bulk opal concentration and to the organic carbon content of the sediment. The abundance of radiolarians and other siliceous microfossils within the lower Oligocene to lower Miocene is interpreted as reflecting changes in surface-water productivity. With only a few exceptions, no stratigraphic indicator species were seen in samples from either Site 646 or Site 647. The absence of both tropical/subtropical and Norwegian-Greenland Sea stratigraphic forms is due to the dominance of subarctic North Atlantic taxa in Leg 105 assemblages. The early Oligocene and early Miocene assemblages recovered at Site 647 are of particular interest, as very little material of these ages has previously been recovered from the subarctic North Atlantic region, and virtually no descriptive work has been conducted on the more endemic components of the radiolarian assemblages from these time intervals. Thus, this report concentrates on providing, at least in part, the first comprehensive documentation of early Oligocene and early Miocene radiolarians from the subarctic North Atlantic, with emphasis on basic descriptions, measurements, and photographic documentation. However, synonymic work and formal designation of new species names has been deferred until additional material from other regions can be examined. The sole exception is the emendation of Theocalyptra tetracantha Bjorklund and Kellogg 1972 to Cycladophora tetracantha n. comb.

Tintinnids of the western Arabian Sea

The results of an investigation of tintinnids from the western Arabian Sea are described. A total of 134 closing-net samples was obtained from 22 stations of the German "Meteor" expedition 1964/1965. Distribution charts of the dominant species of tintinnids from the study area are presented as well as a list of the world-wide distribution of these species as derived from the literature. Tintinnids were most abundant in the surface waters. The layer from 0 - 25 m yielded a maximum 94.3% and a minimum of 61.3% of the tintinnids present from 0 - 175 m; the mean was 80%. There was no significant difference in the vertical distribution between day and night stations nor was there any indication of the influence of the thermocline upon vertical distribution of tintinnids. TS-diagrams show different water types in the western Arabian Sea. Temperatur-salinity-tintinnid -diagrams indicate regional patterns in the distribution of various species of tintinnids. Some tintinnids can be used as indicator species: Climacocylis scalaria, Parundella lohmanni and Amphorella amphora were typical for the Somali Current whereas Rhabdonella apophysata and Branditella palliata indicated the presence of East African Coastal Current water. The concentration of tintinnids in the upper 25 m raged between 4,800 and 39,300 individuals/m**3 (mean 19,000/m**3). Plasma volume of tintinnids was calculated to permit comparison of different links in the food chain. There was a mean of 51 mm**3/m**2 in the upper layer, equivalent to a concentration of 2 mm**3/m**3. Carbon values were computed from the plasma volume of tintinnids, phytoplankton and larger zooplankton. The ratio of phytoplankton plus microzooplankton carbon to large zooplankton carbon was 1 : 0.8 in the Somali Current, 1 : 0.4 in the East African Coastal Current and 1 : 1.2 in the mixing zone of these current systems. Tintinnids are one of the first links in the food chain. It is very likely that a part of the organic detritus and of the nanoplankton is transfered to large herbivores or omnivores via tintinnids and other protozoans. This mechanism might be especially effective during seasons when large phytoplankters are not available in the ocean.

Stable isotope ratios on 3 sediment cores from Tagus prodelta off Lisbon

A high-resolution sedimentary sequence recovered from the Tagus prodelta has been studied with the objective to reconstruct multi-decadal to centennial-scale climate variability on the western Iberian Margin and to discuss the observations in a wider oceanographic and climatic context. Between ca. 100 BC and AD 400 the foraminiferal fauna and high abundance of Globorotalia inflata indicate advection of subtropical waters via the Azores Current and the winter-time warm Portugal Coastal Current. Between ca. AD 400 and 1350, encompassing the Medieval Climate Anomaly (MCA), enhanced upwelling is indicated by the planktonic foraminiferal fauna, in particular by the high abundance of upwelling indicator species Globigerina bulloides. Relatively light d18O values and high sea surface temperature (SST) (reconstructed from foraminiferal assemblages) point to upwelling of subtropical Eastern North Atlantic Central Water. Between ca. AD 1350 and 1750, i.e. most of the Little Ice Age, relatively heavy d18O values and low reconstructed SST, as well as high abundances of Neogloboquadrina incompta, indicate the advection of cold subpolar waters to the area and a southward deflection of the subpolar front in the North Atlantic, as well as changes in the mode of the North Atlantic Oscillation. In addition, the assemblage composition together with the other proxy data reveals less upwelling and stronger river input than during the MCA. Stronger Azores Current influence on the Iberian Margin and strong anthropogenic effect on the climate after AD 1750 is indicated by the foraminiferal fauna. The foraminiferal assemblage shows a significant change in surface water conditions at ca. AD 1900, including enhanced river runoff, a rapid increase in temperature and increased influence of the Azores Current. The Tagus record displays a high degree of similarity to other North Atlantic records, indicating that the site is influenced by atmospheric-oceanic processes operating throughout the North Atlantic, as well as by local changes.

Eemian temperature reconstruction for European sites

Botanical data are widely used as terrestrial proxy data for climate reconstructions. Using a newly established method based on probability density functions (pdf-method), the temperature development throughout the last interglacial, the Eemian, is reconstructed for the two German sites Bispingen and Grobern and the French site La Grande Pile. The results are compared with previous reconstructions using other methods. After a steep increase in January as well as July temperatures in the early phase of the interglacial, the reconstructed most probable climate appears to be slightly warmer than today. While the temperature is reconstructed as relatively stable throughout the Eemian, a certain tendency towards cooler January temperatures is evident. January temperatures decreased from approx. 2-3° C in the early part to approx. -3° C in the later part at Bispingen, and from approx. 2° C to approx. -1° C at Grobern and La Grande Pile. A major drop to about -8° C marks the very end of the interglacial at all three sites. While these results agree well with other proxy data and former reconstructions based on the indicator species method, the results differ significantly from reconstructions based on the modern pollen analogue technique ("pollen transfer functions"). The lack of modern analogues is assumed to be the main reason for the discrepancies. It is concluded that any reconstruction method needs to be evaluated carefully in this respect if used for periods lacking modern analogous plant communities.

Macrofauna biodiversity from ECO2 water-formation experiment

The injection of gas into sub-seabed aquifers may lead to the displacement of hypoxic and hypersaline fluids (reservoir formation water) major environmental risk. To investigate this risk, the impact of formation water release on the macrofaunal community in a mesocosm experiment at Solbergstrand was conducted. 20 boxcores were exposed to 4 treatments (high salinity, hypoxic, mixed and tidal) during two weeks. The abundance of macrofauna was quantified for each treatment and richness, eveness and biodiversity indices calculated. The data are reported in this dataset.

ELPA (European Leaf Physiognomic Approach): Grid data set of the leaf physiognomic composition of the extant European hardwood vegetation

Physiognomic traits of plant leaves such as size, shape or margin are decisively affected by the prevailing environmental conditions of the plant habitat. On the other hand, if a relationship between environment and leaf physiognomy can be shown to exist, vegetation represents a proxy for environmental conditions. This study investigates the relationship between physiognomic traits of leaves from European hardwood vegetation and environmental parameters in order to create a calibration dataset based on high resolution grid cell data. The leaf data are obtained from synthetic chorologic floras, the environmental data comprise climatic and ecologic data. The high resolution of the data allows for a detailed analysis of the spatial dependencies between the investigated parameters. The comparison of environmental parameters and leaf physiognomic characters reveals a clear correlation between temperature related parameters (e.g. mean annual temperature or ground frost frequency) and the expression of leaf characters (e.g. the type of leaf margin or the base of the lamina). Precipitation related parameters (e.g. mean annual precipitation), however, show no correlation with the leaf physiognomic composition of the vegetation. On the basis of these results, transfer functions for several environmental parameters are calculated from the leaf physiognomic composition of the extant vegetation. In a next step, a cluster analysis is applied to the dataset in order to identify "leaf physiognomic communities". Several of these are distinguished, characterised and subsequently used for vegetation classification. Concerning the leaf physiognomic diversity there are precise differences between each of these "leaf physiognomic classes". There is a clear increase of leaf physiognomic diversity with increasing variability of the environmental parameters: Northern vegetation types are characterised by a more or less homogeneous leaf physiognomic composition whereas southern vegetation types like the Mediterranean vegetation show a considerable higher leaf physiognomic diversity. Finally, the transfer functions are used to estimate palaeo-environmental parameters of three fossil European leaf assemblages from Late Oligocene and Middle Miocene. The results are compared with results obtained from other palaeo-environmental reconstructing methods. The estimates based on a direct linear ordination seem to be the most realistic ones, as they are highly consistent with the Coexistence Approach.

Sedimentology, geochemistry and ostracodes from Lake Ohrid

Ancient Lake Ohrid, located in the southern Balkan Peninsula in Macedonia and Albania is characterized by a high degree of endemism and it is considered to be the oldest lake in Europe. But its exact age (between one and ten million years) and also its origin are so far not known. To unravel these uncertainties an ICDP (International Continental Scientific Drilling Program) drilling project (Scientific Collaboration On Past Speciation Conditions in Ohrid (SCOPSCO)), started in April 2013. In addition to the investigations about the age and origin, other paleolimnological studies, e.g., the reconstruction of past climate and of past lake level changes, should be performed with the drilled cores. Used proxies in such paleolimnological studies are, e.g., ostracodes because they respond sensitively to environmental changes but an accurate knowledge of their preferences and tolerances to specific environmental conditions is necessary for this purpose. So far, this knowledge about the, mostly endemic, Ohrid ostracodes was limited. Thus, within the framework of this thesis, ostracodes and a multiplicity of environmental data were collected in Lake Ohrid and its adjacent waters during four field campaigns. In a total of 47 ostracode species could be detected in the entire study area and 32 of them were found alive in Lake Ohrid. Multivariate statistic identified that water depth, salinity, conductivity, pH, and dissolved oxygen were the main determining factors for ostracode distribution in the entire study area. In Lake Ohrid, the distribution was mainly controlled by water depth, water temperature, and pH. Some ostracodes were identified as strong indicator species for important environmental variables, e.g., water temperature and water depth. A distinctive feature of Lake Ohrid was the finding of the ostracode genus Amnicythere whose species normally inhabit oligo-(meso-)haline waters and this could point to a marine origin of the lake. So far, the specialized endemic ostracodes show the highest abundances and the greatest spatial distribution in Lake Ohrid but during the sampling eight widespread species were found for the first time in the lake. They inhabited mainly the northern part of the lake, where two cities are located and industry and agriculture play a major role, and they were limited to water depths above 50 m and this could be an evidence for an increasing anthropogenic pressure because widespread ostracode species often replace endemic species. To unravel the human impact on Lake Ohrid during the last decades short sediment cores were taken and the multi-proxy study indicated that the lake productivity between the early 1920s and the late 1980s was relatively low. Diatom assemblages indicate a rising productivity in the southern part of Lake Ohrid since the mid 1970s and geochemical proxies and ostracodes point to an increasing productivity since the late 1980s in the southern and in the northern part. A slight increase in the productivity continued until 2009. Noticeable is the fact that since the early 1990s, the increasing productivity and the increasing concentrations of heavy metals correspond to a decreasing number of ostracodes in the northern part of Lake Ohrid. Perhaps, this indicates that living conditions in this lake part became less favorable for the mostly endemic ostracode species. Furthermore, the sediment samples from the cores show relatively high concentrations of arsenic, iron, and nickel. Fluctuations in ostracode assemblages from three longer sediment cores, the longest spans approximately 136 ka, taken in Lake Ohrid, correspond to fluctuations in the productivity, in the carbonate content, of the lake level, and of climate changes. Between the marine isotope stage (MIS) 6 and MIS 2 the number of ostracode valves is very low or the valves were completely absent. This corresponds to a low lake productivity, a low carbonate content, and a low lake level. At the onset of the Holocene, the number of valves increased markedly and this correlates with an increased productivity and carbonate content and a warmer climate. But during the Little Ice Age (LIA), the number of valves dropped again and species which prefer warmer waters disappeared completely. This drop corresponds also to a low productivity. After the LIA, the number of species increased again but since 1895 AD a strong and abrupt decrease is visible. A reason for this could be an increase in the heavy metal concentrations.

(Table B2) Relative abundances of diatom species used in the Canoco analysis at sediment surface samples from the Chinese inshore waters

Canonical correspondence analysis has been used to analyze and to visualize the relationships between the main species and selected environmental variables in a study of diatoms from surface sediment samples in Chinese inshore waters. The result shows that the diatom distribution in Chinese inshore waters is closely correlated with the environmental variables and that the measured environmental variables account for the major changes of the diatom composition. Winter sea-surface temperature (WST), winter sea-surface salinity (WSS), water depth and summer sea-surface salinity (SSS) play an important role for the diatom distribution. Among the environmental factors, winter sea-surface temperature is the most important, controlling the distribution of diatoms in the surface sediments in Chinese inshore waters, and therefore, it may be potentially reconstructed in palaeoceanographic studies. Three diatom assemblages are distinguished, representing environments with different hydrological characteristics. The temperate-water diatom assemblage may be used as an indicator of the coastal circulation system of Bohai Sea and Yellow Sea. While the warm-temperate water diatom assemblage is closely related to Shanghai-Zhejiang-Fujian coastal currents and Northern Bay coastal currents of South China Sea. The deep water diatom assemblage is a response to that the waters are less controlled by coastal currents, but are more influenced by open sea currents, such as Kuroshio.

Phenotypic evolution in a venerid bivalve species lineage from the late Middle Miocene Central Paratethys Sea: a multi-approach morphometric analysis

A morphometric analysis was performed for the late Middle Miocene bivalve species lineage of Polititapes tricuspis (Eichwald, 1829) (Veneridae: Tapetini). Specimens from various localities grouped into two stratigraphically successive biozones, i.e. the upper Ervilia Zone and the Sarmatimactra Zone, were investigated using a multi-method approach. A Generalized Procrustes Analysis was computed for fifteen landmarks, covering characteristics of the hinge, muscle scars, and pallial line. The shell outline was separately quantified by applying the Fast Fourier Transform, which redraws the outline by fitting in a combination of trigonometric curves. Shell size was calculated as centroid size from the landmark configuration. Shell thickness, as not covered by either analysis, was additionally measured at the centroid. The analyses showed significant phenotypic differentiation between specimens from the two biozones. The bivalves become distinctly larger and thicker over geological time and develop circular shells with stronger cardinal teeth and a deeper pallial sinus. Data on the paleoenvironmental changes in the late Middle Miocene Central Paratethys Sea suggest the phenotypic shifts to be functional adaptations. The typical habitats for Polititapes changed to extensive, very shallow shores exposed to high wave action and tidal activity. Caused by the growing need for higher mechanical stability, the bivalves produced larger and thicker shells with stronger cardinal teeth. The latter are additionally shifted towards the hinge center to compensate for the lacking lateral teeth and improve stability. The deepening pallial sinus is related to a deeper burrowing habit, which is considered to impede being washed out in the new high-energy settings.