Prolonged and flexible primary moult overlaps extensively with breeding in beach-nesting Hooded Plovers Thinornis rubricollis

We present the first report of complete overlap of breeding and moult in a shorebird. In southeastern Australia, Hooded Plovers Thinornis rubricollis spend their entire lives on oceanic beaches, where they exhibit biparental care. Population moult encompassed the 6-month breeding season. Moult timing was estimated using the Underhill-Zucchini method for Type 2 data with a power transformation to accommodate sexual differences in rates of moult progression in the early and late stages of moult. Average moult durations were long in females (170.3 ± 14.2 days), and even longer in males (210.3 ± 13.5 days). Breeding status was known for most birds in our samples, and many active breeders (especially males) were also growing primaries. Females delayed the onset of primary moult but were able to increase the speed of moult and continue breeding, completing moult at about the same time as males. The mechanism by which this was achieved appeared to be flexibility in moult sequence. All moult formulae fell on one of two linked moult sequences, one faster than the other. The slower sequence had fewer feathers growing concurrently and also had formulae indicating suspended moults. Switching between sequences via common formulae is possible at many points during the moult cycle, and three of 12 recaptures were confirmed to have switched sequences in the same moult season. Hooded Plovers thus have a prolonged primary moult with the flexibility to change their rate of moult; this may facilitate high levels of replacement clutches that are associated with passive nest defence and low reproductive success. © 2014 British Ornithologists' Union.

Storm versus calm : Variation in fauna associated with drifting macrophytes in sandy beach surf zones

New insights about nearshore dynamics came from studying the effects of regular storms in South Australia on drifting marine macrophytes, consequent wrack accumulation and associated fauna in beach surf zones across three different regions. This study examined whether the influence of storms may be more pronounced in sheltered coastal waters compared to more exposed coastlines where biota could have adaptations to persist in larger swell conditions. There were obvious regional differences for wrack species richness, abundances and assemblages that matched the attached floral subtidal landscape in each region. Consequently, invertebrates also differed amongst regions, which highlight the close affinity that some invertebrates have with drifting macrophytes. Fish were not so closely aligned to the regional patterns identified for wrack or invertebrates suggesting that many fish are using wrack accumulations as habitat but, being highly mobile, they may actively and constantly move into, out of and within these habitat features. Well-known beach-type models focused upon beach morphology may be more pertinent to the ecology of the surf zones offshore than previously thought, being the most consistent indicator of wrack accumulations and their fauna. This new evidence on the ecology of nearshore waters during storm versus calm weather in multiple regions and the subsequent influence on wrack-fauna associations in sandy-beach surf zones are important for future beach management, particularly when and where large wrack accumulations occur.

Conservation gone to the dogs : when canids rule the beach in small coastal reserves

On most developed coastlines, dunes backing ocean beaches constitute an urbanised landscape mosaic containing remnant pockets of small conservation areas. Urbanised beaches are also prime sites for domestic dogs, known to be environmentally harmful in many other settings. It is unknown, however, whether small, protected parcels of dune are adequate for biological conservation and whether dogs compromise their functional conservation objectives. Here we examine, for two small (2 km ocean boundary) reserves in Eastern Australia abutting an urban area, whether such small reserves can continue to function as effective conservation instruments on ocean beaches, using scavenger community composition and efficiency to assess ecosystem function. Two non-native species of canids—domestic dogs (Canis lupus familiaris) and red foxes (Vulpes vulpes)—were ubiquitous and numerous inside conservation areas, to the point of having become the most abundant vertebrate scavengers at the beach-dune interface, outcompeting native scavengers for wave-cast carrion. Dogs and foxes have effectively supplanted raptors, normally abundant on non-urban beaches in the region, and other avian scavengers, as the principal consumers of animal carcasses both inside the declared reserves and at the urban beach. Whilst the ecological threats posed by foxes are widely and intensively addressed in Australia in the form of fox-control programs, dog controls are less common and stringent. Our data emphasize, however, that managing domestic dogs may be required to the same extent in order to maintain key forms and functions in coastal reserves situated close to urban areas.

Affective energies : Sensory bodies on the beach in Darwin, Australia

Emerging debates on anti-racism within white majority cultures centre emotion and affect to explore the visceral nature of racialised encounters that unfold in public spaces of the city. This paper builds on such understandings by conceptualising whiteness as a force that exerts affective pressures on bodies of colour who are hypervisible in public spaces. I show that these pressures have the potential to wound, numb and immobilise bodies affecting what they can do or what they can become. This paper argues, however, that affective energies from human and non-human sources are productive forces that are also sensed in public spaces such as the suburban beach. These energies entangle sensuous bodies with the richness of a more-than-human world and have the potential to offer new insights into exploring how racially differentiated bodies live with difference. The paper draws on ethnographic research conducted in Darwin, a tropical north Australian city at the centre of politicised public debates on asylum seeker policy, migrant integration and Indigenous wellbeing. My attention to affective pressures and affective energies contributes to understanding how bodies with complex histories and geographies of racialisation can inhabit a world of becoming. © 2014 Elsevier Ltd. All rights reserved.

Golden opportunities: a horizon scan to expand sandy beach ecology

Robust ecological paradigms and theories should, ideally, hold across several ecosystems. Yet, limited testing of generalities has occurred in some habitats despite these habitats offering unique features to make them good model systems for experiments. We contend this is the case for the ocean-exposed sandy beaches. Beaches have several distinctive traits, including extreme malleability of habitats, strong environmental control of biota, intense cross-boundary exchanges, and food webs highly reliant on imported subsidies. Here we sketch broad topical themes and theoretical concepts of general ecology that are particularly well-suited for ecological studies on sandy shores. These span a broad range: the historical legacies and species traits that determine community assemblages; food-web architectures; novel ecosystems; landscape and spatial ecology and animal movements; invasive species dynamics; ecology of disturbances; ecological thresholds and ecosystem resilience; and habitat restoration and recovery. Collectively, these concepts have the potential to shape the outlook for beach ecology and they should also encourage marine ecologists to embrace, via cross-disciplinary ecological research, exposed sandy beach systems that link the oceans with the land.

Stakeholder knowledge of threatened coastal species; the case of beach-goers and the Hooded Plover Thinornis rubricollis

One way of measuring pre-existing knowledge of a threatened species and its circumstances is to measure the degree of surprise expressed by stakeholders in relation to factual statements regarding the species. Beach-goers (n = 684) were surveyed in regard to their knowledge of the beach-dwelling, threatened, Hooded Plover Thinornis rubricollis, a coastal obligate in south eastern Australia. Principle components analysis revealed that respondents’ degree of knowledge could be categorized as involving ‘chick (flightless young) ecology’ and ‘human impacts’ (threatening processes). Respondents were more surprised by aspects of chick ecology than by threatening processes (F1,514 = 460.446, p < 0.001). Prior knowledge of the species was associated with less surprise at factual statements. Therefore, priorities for further education should focus on linking threats with chick ecology, particularly because an understanding that chicks are not stationary within fenced areas is critical to the interpretation and effectiveness of current signage used to mitigate human impacts.

Breeding habitat selection in an obligate beach bird: a test of the food resource hypothesis

The food resource hypothesis of breeding habitat selection in beach-nesting birds suggests that birds breed at sites with more prey to meet the increased energetic requirements associated with breeding. We compare prey resources using pitfall traps and core samples at breeding sites and absence sites of the eastern population of hooded plover, Thinornis rubricollis rubricollis, which, in this part of its range, is a threatened obligate beach bird. Breeding sites had higher abundances, equivalent species richness, and different assemblages of invertebrate prey compared with absence sites. Assemblages at breeding sites were characterised by more isopods, and fewer beetles of the family Phycosecidae. Breeding habitat selection by plovers appears to be associated with selection for sites with more food, and any process that degrades food resources at a site (e.g. kelp harvesting or marine pollution events) may reduce the likelihood of occupancy of that site by breeding birds.

The role of the libero in volleyball as a paradoxical influence on the game: logical debate and the proposal for a rule change

The purpose of this paper is to bring into focus philosophical and logical concerns relating to specific rules that govern the role of the libero in volleyball. The authors contest that the introduction of the position of the libero in volleyball may have actually undermined the continuing logic of the rules of the game. In addressing these questions the authors offer theoretically based rule-change proposals for the sport of volleyball that could be used to inform future training methods or development programmes at both the participatory and elite levels. Discussion may also inform the development of new tools of analysis (e.g. computer software) or the creation of a new breadth of match simulation resources to enhance athlete performance. These tools and devices could then be presented to volleyball coaches in an effort to resolve the issues that have come to light and further inform the philosophical and logical debates about the role of the libero in volleyball.

"I can sit on the beach and punt through my mobile phone" : the influence of physical and online environments on the gambling risk behaviours of young men.

Gambling is rapidly emerging as an important public health issue, with gambling products causing considerable health and social harms to individuals, families and communities. Whilst researchers have raised concerns about online wagering environments, few studies have sought to explore how factors within different gambling environments (both online and land-based) may be influencing the wagering, and more broadly the gambling risk behaviours of young men. Using semi-structured interviews with 50 Australian men (20-37 years) who gambled on sport, we explored the ways in which online and land-based environments may be risk-promoting settings for gambling. This included the appeal factors associated with gambling in these environments, factors that encouraged individuals to gamble, and factors that encouraged individuals to engage in different, and more harmful types of gambling. Interviews were conducted over the course of a year (April 2015 - April 2016). We identified a number of situational and structural factors that promoted risky gambling environments for young men. In the online environment, gambling products had become exceedingly easy to access through mobile technologies, with young men subscribing to multiple accounts to access industry promotions. The intangibility of money within online environments impacted upon risk perceptions. In land-based environments, the social rituals associated with peer group behaviour and sport influenced risky patterns of gambling. The presence of both gambling and alcohol in pub environments led individuals to gamble more than they normally would, and on products that they would not normally gamble on. Land-based venues also facilitated access to multiple forms of gambling under the one roof. We identified a number of factors in both land and online environments that when combined, created risk-promoting settings for gambling among young men. By exploring these contextual conditions that give rise to gambling harm, we are better able to advocate for effective public health responses in creating environments that prevent harmful gambling.

Regional drivers of clutch loss reveal important trade-offs for beach-nesting birds

Coastal birds are critical ecosystem constituents on sandy shores, yet are threatened by depressed reproductive success resulting from direct and indirect anthropogenic and natural pressures. Few studies examine clutch fate across the wide range of environments experienced by birds; instead, most focus at the small site scale. We examine survival of model shorebird clutches as an index of true clutch survival at a regional scale (∼200 km), encompassing a variety of geomorphologies, predator communities, and human use regimes in southeast Queensland, Australia. Of the 132 model nests deployed and monitored with cameras, 45 (34%) survived the experimental exposure period. Thirty-five (27%) were lost to flooding, 32 (24%) were depredated, nine (7%) buried by sand, seven (5%) destroyed by people, three (2%) failed by unknown causes, and one (1%) was destroyed by a dog. Clutch fate differed substantially among regions, particularly with respect to losses from flooding and predation. ‘Topographic’ exposure was the main driver of mortality of nests placed close to the drift line near the base of dunes, which were lost to waves (particularly during storms) and to a lesser extent depredation. Predators determined the fate of clutches not lost to waves, with the depredation probability largely influenced by region. Depredation probability declined as nests were backed by higher dunes and were placed closer to vegetation. This study emphasizes the scale at which clutch fate and survival varies within a regional context, the prominence of corvids as egg predators, the significant role of flooding as a source of nest loss, and the multiple trade-offs faced by beach-nesting birds and those that manage them.

An obligate beach bird selects sub-, inter- and supra-tidal habitat elements

Few habitat models are available for widespread, obligate, high-energy sandy shore vertebrates, such as the Eastern Hooded Plover Thinornis cucullatus cucullatus. We examined habitat attributes which determined the difference between sites where plovers breed and randomly-selected absence sites (determined from long-term systematic monitoring). A variety of habitat variables were derived from aerial photography and bathymetric and terrestrial Light Detection And Ranging (LiDAR) data. Logistic regression against eight candidate variables, in a model selection framework, revealed considerable support for four variables with respect to explaining the presence of breeding territories. In particular, the amount of unvegetated dune and foredune which was unvegetated, and the amount of intertidal and sub-tidal reef were positively associated with the presence of breeding territories. Thus, plovers apparently select certain habitat in which to breed, involving sub-tidal, intertidal and supra-tidal habitat elements. The model also helps explain the virtual absence of breeding plovers from long sections of superficially suitable habitat, such as the fourth longest continuous beach in the world.

Prey selection, size, and breakage differences in Turbo undulatus opercula found within Pacific Gull (Larus pacificus) middens compared to Aboriginal middens and natural beach deposits, southeast Australia

Qualitative discrimination criteria are employed commonly to distinguish cultural shell middens from natural shell deposits. Quantitative discrimination criteria remain less developed beyond an assumption that natural shell beds tend to contain a wider range of shell sizes compared to cultural shell middens. This study further tests this assumption and provides the first comparative quantitative analysis of shell sizes from cultural middens, bird middens, and beach shell beds. Size distributions of opercula of the marine gastropod Turbo undulatus within two modern Pacific Gull (Larus pacificus) middens are compared with two Aboriginal middens (early and late Holocene) and two modern beach deposits from southeast Australia. Results reveal statistically significant differences between bird middens and other types of shell deposits, and that opercula size distributions are useful to distinguish Aboriginal middens from bird middens but not from beach deposits. Supplementary qualitative analysis of taphonomic alteration of opercula reveal similar opercula breakage patterns in human and bird middens, and further support previously recognised criteria to distinguished beach deposits (water rolling and bioerosion) and human middens (burning). Although Pacific Gulls are geographically restricted to southern Australia, the known capacity of gulls (Larus spp.) in other coastal contexts around the world to accumulate shell deposits indicates the broader methodological relevance of our study.

Selecting umbrella species for conservation: a test of habitat models and niche overlap for beach-nesting birds

Umbrella species are rarely selected systematically from a range of candidate species. On sandy beaches, birds that nest on the upper beach or in dunes are threatened globally and hence are prime candidates for conservation intervention and putative umbrella species status. Here we use a maximum-likelihood, multi-species distribution modeling approach to select an appropriate conservation umbrella from a group of candidate species occupying similar habitats. We identify overlap in spatial extent and niche characteristics among four beach-nesting bird species of conservation concern, American oystercatchers (Haematopus palliatus), black skimmers (Rynchops niger), least terns (Sterna antillarum) and piping plovers (Charadrius melodus), across their entire breeding range in New Jersey, USA. We quantify the benefit and efficiency of using each species as a candidate umbrella on the remaining group. Piping plover nesting habitat encompassed 86% of the least tern habitat but only 15% and 13% of the black skimmer and American oystercatcher habitat, respectively. However, plovers co-occur with all three species across 66% of their total nesting habitat extent (~ 649 ha), suggesting their value as an umbrella at the local scale. American oystercatcher nesting habitat covers 100%, 99% and 47% of piping plover, least tern and black skimmer habitat, making this species more appropriate conservation umbrellas at a regional scale. Our results demonstrate that the choice of umbrella species requires explicit consideration of spatial scale and an understanding of the habitat attributes that an umbrella species represents and to which extent it encompasses other species of conservation interest. Notwithstanding the attractiveness of the umbrella species concept, local conservation interventions especially for breeding individuals in small populations may still be needed.