Genome-wide evidence for speciation with gene flow in Heliconius butterflies

Most speciation events probably occur gradually, without complete and immediate reproductive isolation, but the full extent of gene flow between diverging species has rarely been characterized on a genome-wide scale. Documenting the extent and timing of admixture between diverging species can clarify the role of geographic isolation in speciation. Here we use new methodology to quantify admixture at different stages of divergence in Heliconius butterflies, based on whole-genome sequences of 31 individuals. Comparisons between sympatric and allopatric populations of H. melpomene, H. cydno, and H. timareta revealed a genome-wide trend of increased shared variation in sympatry, indicative of pervasive interspecific gene flow. Up to 40% of 100-kb genomic windows clustered by geography rather than by species, demonstrating that a very substantial fraction of the genome has been shared between sympatric species. Analyses of genetic variation shared over different time intervals suggested that admixture between these species has continued since early in speciation. Alleles shared between species during recent time intervals displayed higher levels of linkage disequilibrium than those shared over longer time intervals, suggesting that this admixture took place at multiple points during divergence and is probably ongoing. The signal of admixture was significantly reduced around loci controlling divergent wing patterns, as well as throughout the Z chromosome, consistent with strong selection for Müllerian mimicry and with known Z-linked hybrid incompatibility. Overall these results show that species divergence can occur in the face of persistent and genome-wide admixture over long periods of time.

Soft Power del Vaticano en el reconocimiento de Palestina como Estado

Desde 1945, el inicio de la cuestión palestina, el país ha sufrido una serie de transformaciones e intervenciones en su soberanía y en legitimidad, las cuales siguen en discusión actualmente; y han dado como resultado el estudio del reconocimiento de Palestina como Estado. Es pertinente hacer un acercamiento a este tema desde la variable religiosa (Vaticano), teniendo en cuenta el gran número de lugares sagrados que hay en la región de Palestina y la variable religiosa del conflicto Palestino-israelí. Desde la mirada del Soft Power, concepto que ha venido tomando fuerza en la academia desde 1980, podemos acercarnos al objeto de estudio y dilucidar las injerencias de este actor. Mediante una aproximación cualitativa, que conlleva análisis del discurso, análisis histórico y análisis de política exterior, se logrará obtener el resultado esperado, que es entender de qué manera el Vaticano ejerce influencia (ideológica y cultural) en el Sistema Internacional, en el asunto del reconocimiento de Palestina como Estado.