Plant Size as Determinant of Species Richness of Herbivores, Natural Enemies and Pollinators across 21 Brassicaceae Species

Large plants are often more conspicuous and more attractive for associated animals than small plants, e.g. due to their wider range of resources. Therefore, plant size can positively affect species richness of associated animals, as shown for single groups of herbivores, but studies usually consider intraspecific size differences of plants in unstandardised environments. As comprehensive tests of interspecific plant size differences under standardised conditions are missing so far, we investigated effects of plant size on species richness of all associated arthropods using a common garden experiment with 21 Brassicaceae species covering a broad interspecific plant size gradient from 10 to 130 cm height. We recorded plant associated ecto-and endophagous herbivores, their natural enemies and pollinators on and in each aboveground plant organ, i.e. flowers, fruits, leaves and stems. Plant size (measured as height from the ground), the number of different plant organ entities and their biomass were assessed. Increasing plant size led to increased species richness of associated herbivores, natural enemies and pollinating insects. This pattern was found for ectophagous and endophagous herbivores, their natural enemies, as well as for herbivores associated with leaves and fruits and their natural enemies, independently of the additional positive effects of resource availability (i.e. organ biomass or number of entities and, regarding natural enemies, herbivore species richness). We found a lower R-2 for pollinators compared to herbivores and natural enemies, probably caused by the high importance of flower characteristics for pollinator species richness besides plant size. Overall, the increase in plant height from 10 to 130 cm led to a 2.7-fold increase in predicted total arthropod species richness. In conclusion, plant size is a comprehensive driver of species richness of the plant associated arthropods, including pollinators, herbivores and their natural enemies, whether they are endophagous or ectophagous or associated with leaves or fruits.

Age an origin of the "Mexican Onyx" at San Antonio Texcala (Puebla, Mexico)

A great variety of color banded CaC03 is known as „Mexican Onyx“, and is extensively used for ornamental purposes. Within the San Antonio Texcala mining district an area of about 24.000 m2 is covered by a thick travertine crust with a calculated volume of at least 2,8 x 1061. It originates from warm waters that emerge on young fissures and faults. It seems to be likely that the travertine formation is related to the hydrothermal activity of the Transmexican Volcanic Belt and to the seismicity of this zone. The deposit is built up by different travertine varieties. The main lithotypes are: (1) dense crystalline laminated travertine; (2) ray crystal travertine; (3) shrub layer travertine; (4) irregular porous travertine; (5) travertine breccia. Textural variation seems to be related to water temperature/distance from the emergence point, rapid or slow degassing of CO, and bacterial influence. At the moment, a light green banded variety (native sulfur impurities) is mined. It is the facture fill of a morphological prominent fissure-ridge travertine. U/Th data indicate, that the travertine has been deposited since at least 52 + 5 ka.