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Acknowledgments RRP was supported by a PhD-studentship from the University of Valladolid (co-funded by Banco Santander, RR 30/04/2014). Financial support was provided by ECOCYCLES (BIODIVERSA 2008, Era-net European project, EUI2008-03658 and NERC NE/G002045/1 to XL) and ECOVOLE projects (CGL2012-35348; Ministerio de Economía y Competitividad of Spain). The article also contributes to project ECOTULA (CGL2015-66962-C2-1-R). We held all the necessary licenses and permits for conducting this work (JJLL, FM and RRP held animal experimentation permits of level B for Spain, and a capture permit was provided by the Consejería de Fomento y Medio Ambiente, Junta de Castilla y León (Expte: EP/CYL/665/2014)). We thank two anonymous reviewers for providing and constructive comments to improve the manuscript.