Vertical and horizontal movements of southern bluefin tuna (Thunnus maccoyii) in the Great Australian Bight observed with ultrasonic telemetry

The vertical and horizontal movements of southern bluefin tuna (SBT), Thunnus maccoyii, in the Great Australian Bight were investigated by ultrasonic telemetry. Between 1992 and 1994, sixteen tuna were tracked for up to 49 h with depth or combined temperature-depth transmitting tags. The average swimming speeds (measured over the ground) over entire tracks ranged from 0.5 to 1.4 m/s or 0.5 to 1.4 body lengths/s. The highest sustained swimming speed recorded was 2.5 m/s for 18 hours. Horizontal movements were often associated with topographical features such as lumps, reefs, islands and the shelf break. They spent long periods of time at the surface during the day (nearly 30%), which would facilitate abundance estimation by aerial survey. At night, they tended to remain just below the surface, but many remained in the upper 10 m throughout the night. SBT were often observed at the thermocline interface or at the surface while travelling. A characteristic feature of many tracks was sudden dives before dawn and after sunset during twilight, followed by a gradual return to their original depth. It is suggested that this is a behavior evolved to locate the scattering layer and its associated prey when SBT are in waters of sufficient depth. SBT maintained a difference between stomach and ambient temperature of up to 9°C.

Decadal variation in the trans-Pacific migration of northern bluefin tuna (Thunnus thynnus) coherent with climate-induced change in prey abundance [abstract]

EXTRACT (SEE PDF FOR FULL ABSTRACT): Indices of the relative abundance of bluefin tuna in the western and eastern Pacific show decadal variation in the proportion of bluefin making trans-Pacific migrations out of the western Pacific.

World meeting on stock assessment of bluefin tunas: strengths and weaknesses

The Inter-American Tropical Tuna Commission (IATTC) hosted a meeting, sponsored jointly by the IATTC and the Australian Fisheries Service, to discuss and report on the strengths and weaknesses of stock assessment techniques used on bluefin tuna stocks in the Pacific, Indian, and Atlantic Oceans and the Mediterranean Sea. The meeting was held in La Jolla, California, on Mat 25-31, 1990.

Symposium on World tuna fisheries: commemorating the 50th anniversary of the establishment of the Inter-American Tropical Tuna Commission

The Inter-American Tropical Tuna Commission (lA TTC) came into existence in 1950, after its convention, signed by representatives ofCosta Rica and the United States in 1949, was ratified. It was the first international tuna organization, and only the third international fisheries organization, whose staff has had the responsibility for performing scientific research, the others being the International Pacific Halibut Commission, established in 1923, and the International Pacific Salmon Fisheries Commission, established in 1937. The current members of the IATTC are Costa Rica, Ecuador, EI Salvador, France, Guatemala, Japan, Mexico, Nicaragua, Panama, the United States, Vanuatu, and Venezuela. The first Director ofthe IATTC was Dr. Milner B. Schaefer, who was in that position from 1950 to 1963. He was followed by Dr. John L. Kask (1963-1969), Dr. James Joseph (1969-1999), and Dr. Robin L. Allen (1999-present). The success ofthe IATIC showed that it was possible to carry out research and management on an international, high-seas fishery successfully. Since then other international organizations for tuna management, including the International Commission for the Conservation of Atlantic Tunas (1969), the Forum Fisheries Agency (1979), the Commission for the Conservation of Southern Bluefin Tuna (1994), and the Indian Ocean Tuna Commission (1996), were established. Appropriately, the 50th anniversary celebration was held in Costa Rica, one of the two charter members of the IATTC. Persons who have held important positions in international fishery management in various parts ofthe world spoke at the celebration. Their presentations, except for that describing the Indian Ocean Tuna Commission, are reproduced in this volume.

Marine ecosystems of the North Pacific

Key Messages [pdf, 2.5 Mb] Climate Information Gaps Ocean Productivity Information gaps Living Marine Resources Information gaps Climate [pdf, 1.8 Mb] Productivity [pdf, 5.2 Mb] Nutrients Phytoplankton Zooplankton Living Resources [pdf, 10 Mb] Subarctic coastal systems Central oceanic gyres Temperate coastal and oceanic systems Marine mammals The Human Population [pdf, 5 Mb] Contaminants and Habitat Modifications Aquaculture Knowledge Gaps Glossary Ocean and Climate Changes [pdf, 4.1Mb] Highlights Introduction Atmospheric Indices Change in 1998/99 Comparison of Atmospheric Indices Authorship Yellow Sea / East China Sea [pdf, 2.3 Mb] Highlights Background Status and Trends Hydrography Chemistry Plankton Benthos Fish and invertebrates Marine birds and mammals Issues Critical factors causing change Authorship Japan/East Sea [pdf, 3.3 Mb] Highlights Background Status and Trends Hydrography Chemistry Plankton Fish and Invertebrates Marine Birds and Mammals Critical factors causing change Issues Authorship Okhotsk Sea [pdf, 1.7 Mb] Background Status and Trends Climate Hydrography Chemistry Plankton Fish and Invertebrates Marine Birds and Mammals Issues Critical factors causing change Authorship Oyashio / Kuroshio [pdf, 4.5 Mb] Highlights Background Status and Trends Hydrography Plankton Fish and Invertebrates Marine Birds and Mammals Issues Authorship Western Subarctic Gyre [pdf, 4.5 Mb] Highlights Background Status and Trends Hydrography Chemistry Plankton Fish and Invertebrates Marine Birds and Mammals Issues Authorship Bering Sea [pdf, 2.2 Mb] Highlights Background Status and Trends Hydrography Chemistry Plankton Fish and Invertebrates Marine Birds and Mammals Critical Factors Causing Change Issues Authorship Gulf of Alaska [pdf, 2.6 Mb] Highlights Background Status and trends Hydrography Chemistry Plankton Fish and Invertebrates Marine birds and mammals Critical factors causing change Issues Authorship California Current [pdf, 2.7 Mb] Highlights Background Status and Trends Hydrography Chemistry Plankton Fish and Invertebrates Marine Birds and Mammals Critical Factors Causing Change Issues Authorship Gulf of California [pdf, 1.7 Mb] Highlights Background Status and Trends Hydrography Chemistry Plankton Fisheries Marine Birds and Mammals Critical Factors Causing Change Issues Authorship Transition Zone [pdf, 2.5 Mb] Background Status and Trends Hydrography Chemistry Plankton Fish and Invertebrates Marine Birds and Mammals Issues Authorship Tuna [pdf, 1.5 Mb] Highlights Background Pacific bluefin tuna Albacore tuna Status and trends Ecosystem model and climate forcing Authorship Pacific halibut [pdf, 1.1 Mb] Background The Fishery Climate Influences Authorship Pacific salmon [Updated, pdf, 0.4 Mb] Background Status and Trends Washington, Oregon, and California British Columbia Southeast Alaska Central Alaska Western Alaska Russia Japan Authorship References [pdf, 0.5 Mb]

Proximate composition and fatty acid and cholesterol content of 22 species of northwest Atlantic finfish

The moisture, fat, ash, fatty acid profile, and cholesterol content are reported for cooked and raw fillets from 22 species of finfish found in the Northwest Atlantic. All but nine species had 1%or less fat. Ocean perch and a spring sampling of mackerel and wolffiSh had about 2% fat, followed by yellowfin tuna, whiting, silver hake, butterfish, and a summer -sampling of mackerel and wolffish with a range of 3-7% fat. Herring had a range of 5-12% fat representing a winter sampling on the low end and summer sampling on the high end of the range. Bluefin tuna (a summer sampling) contained the most fat with a high of 23% fat. Omega-3 fatty acids were present in excess of omega-6 fatty acids. The fattier fISh supplied the most omega-3 fatty acids per gram of tissue. The mean cholesterol content for all species was 57 ± 16 mg/l00 g raw tissue. Finfish from the Northwest Atlantic would appear to fit into the regime for a healthy heart, being low in fat and cholesterol and rich in omega-3 fatty acids.(PDF file contains 42 pages.)

Fangquoten für die EU-Mitgliedsländer 1998

The fishery ministers of the EU agreed in their yearly pre-Christmas marathon session on 18th and 19th Dec. 1997 on next year’s TACs and country quota. Main decision was the increase of the TAC for North sea cod and a smaller than feared cut of cod TACs in the West of Scotland zone, the Irish Sea and the English Channel. For the first time ministers set country quota for horse mackerel; until 1997 there had been - with the exception for Portugal and Spain - only a common quota. Also for the much disputed sandeel fishery a TAC of 1 mio. t was finally established, as well as for bluefin tuna and swordfish in the Atlantic and Mediterranean. For 1998 the EU member countries dispose of a total catch quota of 4.46 mio. t in EU waters, and together with quota in third country waters of 5.2 mio. t. Of this total quota, 3.16 mio. t are fish for human consumption and 2.1 mio. t fish for industrial purposes.

Evidence of shark predation and scavenging on fishes equipped with pop-up satellite archival tags

Over the past few years, pop-up satellite archival tags (PSATs) have been used to investigate the behavior, movements, thermal biology, and postrelease mortality of a wide range of large, highly migratory species including bluefin tuna (Block et al., 2001), swordfish (Sedberry and Loefer, 2001), blue marlin (Graves et al., 2002), striped marlin (Domeier and Dewar, 2003), and white sharks (Boustany et al., 2002). PSAT tag technology has improved rapidly, and current tag models are capable of collecting, processing, and storing large amounts of information on light level, temperature, and pressure (depth) for a predetermined length of time before the release of these tags from animals. After release, the tags float to the surface, and transmit the stored data to passing satellites of the Argos system.

Growth of yellowfin tuna, Thunnus albacares, in the Eastern Pacific Ocean based on otolith increments

ENGLISH: The growth of yellowfin tuna in the eastern Pacific is described in terms of several measurements taken from the fish and their otoliths (sagittae). Equations are also developed to predict age from the readily available dimensions of fork length and head length. The data for all of these relationships were obtained from a sample of 196 fish collected during 1977 through 1979 from purse seiners fishing north of the equator and east of 137°W. The fork-length range of the sample was 30-170 cm. The number of increments on a sagitta of each fish was used as a direct estimate of its age in days. The correspondence between increments and days has been validated for yellowfin in the length range of 40-110 cm. Circumstantial evidence indicates that the relationship also applies in the intervals of 0-40 cm and 110-170 cm. This circumstancial evidence was derived from: 1) literature on validated increments during early growth for other species, 2) knowledge that structures assumed to be daily increments on yellowfin otoliths have subsequently been validated in the corresponding zone on bluefin otoliths, and 3) a comparison of the growth curve based on increments to others obtained from length frequency modal analysis. Based on this information the age estimates over the entire size range of sampled fish are believed to be accurate. In addition to the general growth and age-predictive relationships, the major conclusions of the study are that: 1) Sexually dimorphic growth exists in terms of fork length, fish weight and the length of the otolith counting path for the entire data set. Examination of the data for 1977 and 1979 also revealed that the fork-length growth of each sex differed within years. 2) For combined sexes there were significant differences among the fork-length growth curves for yellowfin sampled in different years. 3) Yellowfin caught inshore (within 275 miles of the coast) were heavier than those caught offshore for fork lengths between 30 and 110 cm. The situation was reversed for lengths greater than 110 cm. 4) Back-calculated spawning months were distributed uniformly throughout the year in 1974 and 1977, but in 1975-1976 and 1978 spawning activity was apparently concentrated in the latter half of the year. SPANISH: El crecimiento del atún aleta amarilla en el Pacífico oriental se describe en términos de varias medidas obtenidas de peces y otolitos (sagita). Se formularon también ecuaciones para pronosticar la edad, según las dimensiones fácilmente disponibles de la longitud horquilla y longitud de la cabeza. Los datos de todas estas relaciones fueron obtenidos mediante una muestra de 196 peces recolectados desde 1977hasta 1979, en barcos cerqueros que estaban pescando al norte de la línea ecuatorial y al este de los 137°W. El intervalo de la longitud horquilla de la muestra fue de 30-170 cm. Se empleó el número de incrementos en la sagita de cada pez como un estimado directo de la edad en días. Se ha comprobado la relación entre los incrementos y los días en el intervalo de longitud de 40-110 cm del aleta amarilla. La evidencia circunstancial indica que se aplica también la relación a los intervalos de 0-40 cm y 110-170 cm. Esta evidencia circunstancial se dedujo: 1) de las publicaciones sobre incrementos comprobados de otras especies durante el primer crecimiento, 2) del conocimientoque las estructuras que se supone son incrementos diarios en los otolitos del aleta amarilla han sido comprobadas luego en la parte correspondiente de otolitos del aleta azul y 3) por una comparación de la curva de crecimiento, basada en incrementos relacionados a otras curvas obtenidas según el análisis modal frecuencia-talla. Se cree, basados en esta información, que las estimaciones de la edad sobre toda la amplitud de talla de los peces muestreados, es acertada. Además de la relación del crecimiento general y del pronóstico de la edad, las principales conclusiones de este estudio son: 1) En toda la serie de datos existe el crecimiento sexualmente dimórfico en términos de longitud horquilla, peso del pez y longitud del plano de conteo del otolito. El examen de los datos de 1977 y 1979, revelan también que el crecimiento longitud horquilla de cada sexo es diferente en los años. 2) En los sexos combinados hubo diferencias significativas entre las curvas de crecimiento longitud horquilla del aleta amarilla muestreado en diferentes años. 3) El aleta amarilla capturado cerca a la costa (en las primeras 275 millas) fue más pesado que el capturado en las aguas mar afuera, correspondiente a la longitud horquilla entre 30 y 110 cm. La situación fue inversa para tallas de más de 110 cm. 4) En 1974 y 1977, los meses retrocalculados del desove se distribuyeron uniformemente durante el año, pero en 1975-1976 y 1978, la actividad del desove se concentró aparentemente en el último semestre del año. (PDF contains 62 pages.)

U. S. Trade in Tuna for Canning, 1987

U.S. tuna fleet activity, canned tuna processing, ex-vessel, wholesale and retail prices and imports in 1987 are described and compared to their counterparts in previous years. Industry statistics gathered from government agencies and industry contacts are presented in 14 figures and 8 tables. In 1987, U.S. tuna fisheries delivered 253,136 short tons (tons) of tuna to U.S. canneries. Domestic deliveries of albacore (white-meat) tuna were 2,836 tons, down 20 percent from 1986 levels. Domestic deliveries of tropical (light-meat) tuna (bigeye, blackfin, bluefin, skipjack, and yellowfin) were 251,000 tons, up 12 percent. Contract prices for tuna delivered by U. S. vessels to U. S. canneries increased dramatically in 1987. Depending on the size of fish in the delivery, ex-vessel prices of white-meat tuna increased as much as 27 percent, and prices of light-meat tuna increased as much as 47 percent. U. S. cannery receipts of imported and domestically caught raw frozen tuna for canning totaled 532,704 tons in 1987, up 2 percent from 1986 levels. U.S. cannery receipts of white-meat tuna were 104,197 tons, down 10 percent from 1986. Imports made up 97 percent of the total cannery supply. Total 1987 U. S. cannery receipts of raw, frozen light meat tuna were 428,507 tons, up 5 percent from 1986 levels. Imports made up 41 percent of the total cannery supply. The 1987 U.S. pack of canned tuna was 33.6 million standard cases, up 3 percent from 1986. The pack of white-meat tuna was 7.2 million standard cases, down 11 percent from 1986; the pack of light-meat tuna was 26.4 million standard cases, up 7 percent. U. S. imports of canned tuna in 1987 were 10.8 million standard cases, down 11 percent from 1986 levels, the first time in recent years that imports have declined. Per capita consumption of canned tuna in the United States was 3.5 pounds in 1987, down slightly from 1986. The retail composite price was $2.26 per pound, unchanged from 1986.

Estimations, from tagging experiments, of mortality rates and other parameters respecting yellowfin and skipjack tuna

ENGLISH: In this paper, a method of analysis described by Gulland (1963) has been used to estimate the fishing mortality rates of tagged yellowfin and skipjack tuna for specific areas and years. Fishing mortality rates obtained for tagged tunas will also represent those for the entire population from which the tagged fishes were drawn, provided the assumptions used and corrections made for these analyses are valid. Total mortality rates of tagged fishes have also been computed. These are not assumed to be directly equivalent to the total mortality rates of the untagged populations,since tagged fishes are subject to additional types of attrition. These additional sources of mortality are also examined in this study. SPANISH: En el presente trabajo se ha usado un método de análisis descrito por Gulland (1963), para estimar las tasas de mortalidad de pesca de los atunes aleta amarilla y barrilete marcados en áreas y años específicos. Las tasas de mortalidad de pesca obtenidas en atunes marcados representarán también las de toda la población, de la cual fueron extraídos, previendo que las suposiciones usadas y las correcciones hechas para estos análisis sean válidas. Las tasas de mortalidad total de los peces marcados también han sido computadas. No se supone que éstas sean directamente equivalentes a las tasas de mortalidad total de las poblaciones no marcadas, ya que los peces marcados están sujetos también a otros tipos de pérdida. Estas otras causas de mortalidad son examinadas también en el presente estudio.

Size composition of catches of Yellowfin tuna in the Japanese long-line fishery in the eastern tropical Pacific east of 130°W

ENGLISH (pgs. 267-283): In the spring of 1963, the senior author, who is a member of the staff of the Nankai Regional Fisheries Research. Laboratory, Fisheries Agency, Japanese Government, came to the Institute of Marine Resources of the University of California as a visiting investigator, bringing with him catch statistical data from the fishery in the eastern Pacific, which had been collected at the Nankai Regional Fisheries Research Laboratory (NRFRL) through September 1962, in order to conduct studies of these data in collaboration with the junior author, and with investigators of the InterAmerican Tropical Tuna Commission. A general review of the long-line fishery, based on the catch statistics of the commercial fishing fleet has been published by Suda and Schaefer (1965). In this paper we present an analysis of data respecting the size-composition of yellowfin tuna taken on long-line gear throughout the eastern Pacific between 1958 and 1962, and make some comparisons with data on size-composition of yellowfin tuna taken in the near-surface fishery, by bait boats and purse-seiners, in waters adjacent to the American coast. As has been shown by Suda and Schaefer (1965), the long-line fishery in the eastern Pacific is primarily directed toward the capture of bigeye tuna. However, considerable quantities of yellowfin tuna are also taken on this gear, and, in addition, there are substantial catches of albacore and of several species of spearfishes in some parts of the range of this fishery. Information respecting the catch rates of yellowfin tuna, and information respecting the size-composition of the stock of yellowfin tuna being exploited by the long-line fishery, is of particular interes~" because the yellowfin tuna population of the eastern Pacific is also subject to an intensive fishery by baitboats and purse-seiners which capture this species, together with skipjack, near the surface along the coast of the Americas, and around the outlying islands, in the region of California to northern Chile. SPANISH (pgs. 311-329): En la primavera de 1963, el autor principal, quien es miembro del personal del Nankai Regional Fisheries Research Laboratory, Fisheries Agency del gobierno japonés, vino al Institute of Marine Resources de la Universidad de California en calidad de investigador visitante y trajo consigo datos estadísticos de las capturas de la pesquería en el Pacífico oriental, que habían sido recolectados en el Nankai Regional Fisheries Research Laboratory (NRFRL) hasta septiembre de 1962, con el fin de hacer estudios de esos datos en colaboración con el coautor y con investigadores de la Comisión Interamericana del Atún Tropical. Una revisión general de la pesquería con palangre, basada sobre las estadísticas de captura de la flota pesquera comercial, ha sido publicada por Suda y Schaefer (1965). En este trabajo presentamos un análisis de los datos correspondientes a la composición de tamaños del atún aleta amarilla capturado con equipo palangrero en todo el Pacífico oriental, entre 1958 y 1962, y hacemos algunas comparaciones con los datos sobre la composición de tamaños del atún aleta amarilla cogido en la pesquería superficial cercana, por barcos de carnada y rederos en aguas adyacentes a la costa americana. Como ha sido demostrado por Suda y Schaefer (1965) la pesquería con palangre en el Pacífico oriental tiene como principal objeto la captura del atún ojo grande. Sin embargo, considerables cantidades de atún aleta amarilla son capturadas también por este equipo y, además, hay también considerables capturas de albacora y de diversas especies de peces-espada en algunas partes de la región que abarca esta pesquería. La información respecto a las tasas de captura del atún aleta amarilla, y la relativa a la composición de tamaños del stock de esta especie que explota la pesquería con palangre, es de particular interés, a causa de que la población de atún aleta amarilla del Pacífico oriental es también objeto de una pesca intensiva por barcos de carnada y rederos que capturan esta especie, junto con el barrilete, cerca de la superficie a 10 largo de la costa de las Américas y alrededor de las islas mar afuera, en la región desde California hasta el norte de Chile.

Tunas and tuna fisheries of the World: an annotated bibliography, 1930-1953

This bibliography attempts to list, with descriptive annotations and a subject index, important literature published between 1930 and 1953 dealing with the tunas and their fisheries in all parts of the world. It is thus a continuation of Corwin's (1930) work, which extended with similar scope through 1929, and an extension of Shimada's (1951), which was limited to the biology of Pacific tunas. The tunas with which it deals are those fishes customarily so-called in commercial parlance and usually classified in the genera Thunnus, Neothunnus, Parathunnus, Germo, Katsuwonus, Euthynnus and Auxis and their various synonyms. All aspects of the biology of the tunas are dealt with, as are descriptions and histories of all types of tuna fisheries, commercial and exploratory tuna fishing methods and results, fishing gear, catch statistics, and fishery management, but processing technology, economics and marketing, folklore, and purely literary references have been excluded.

Additional information on the length-weight relationship of skipjack tuna from the Eastern Tropical Pacific Ocean

ENGLISH: Results of a study of the length-weight relationships of yellowfin (Neothunnus macropterus) and skipjack (Katsuwonus pelamis) tuna from several fishing areas of the Eastern Tropical Pacific Ocean have been published by Chatwin (1959). In that report, a very low exponential value of 2.6261 was obtained for skipjack from Area 14 (off northern Chile, see Chatwin, Figure 1). It was pointed out, however, that this estimate was based on two samples of fish with a very narrow range of total lengths, not representative of the range in the catch, and that it would be desirable to obtain a further estimate based on a larger range of total lengths. In addition, there proved to be significantly large differences among exponents for the areas sampled, precluding use of a single regression equation for all areas. Two important fishing areas remained unsampled (Areas 10 and 13, see Chatwin, Figure 1), and it appeared desirable to collect length-weight measurement data from them, so that estimating equations would be available for all areas. Subsequent to publication of Chatwin's study, samples of skipjack length-weight measurements were obtained from the desired areas. Estimates derived from these data, and their effects on the previous analysis are presented herein. SPANISH:Los resultados de un estudio sobre las relaciones entre la longitud y el peso del atún aleta amarilla (Neothunnus macropterus) y del barrilete (Katsuwonus pelamis) de las diferentes áreas de pesca en el Pacífico Oriental Tropical ya han sido publicados por Chatwin (1959). En ese informe se obtuvo un valor exponencial muy bajo de 2.626 para el barrilete del Area 14 (frente a la costa norte de Chile, ver Chatwin, Figura 1). Se hizo hincapié, sin embargo, en que esta estimación se basaba en dos muestras de peces con una amplitud muy estrecha de longitudes totales, no representativa de la amplitud en la pesca, y que sería deseable obtener una estimación adicional basada en una amplitud mayor de longitudes totales. Además, se comprobó que habian diferencias significativamente grandes entre los exponentes de las áreas muestreadas lo que impedía el usa de una sola ecuación de regresión para todas las áreas. Se quedaron sin muestrear dos importantes áreas de pesca (Areas 10 y 13, ver Chatwin, Figura 1) y pareció deseable recolectar datos de medidas de longitud y peso de estas áreas, de tal manera que hubiesen disponibles ecuaciones estimadoras para todas las áreas. Después de la publicación del estudio de Chatwin, so obtuvieron muestras de medidas de longitud y peso de barriletes de las áreas deseadas. Las estimaciones derivadas de estos datos y sus efectos sabre el análisis previo se dan en el presente informe.

Distribution of fishing effort and resulting tuna catches from the Eastern Tropical Pacific by quarters of the year, 1951-1958

ENGLISH: This is a data report based on information collected from the logbooks of baitboats and purse-seiners engaged in the fishery for yellowfin (Neothunnus macropterus) and skipjack (Katsulvonus pelamis) tuna in the Eastern Tropical Pacific Ocean from 1951 through 1958. The detailed analysis of these data appears in other research bulletins, some already published and others in preparation. SPANISH: Este es un informe basado en los datos contenidos en la información recogida de los registros de bitácora de los barcos de carnada y rederos que se dedicaron a la pesca de atún aleta amarilla (Neothunnus macropterus) y barrilete (Katsuwonus pelamis) en el Océano Pacífico Oriental Tropical de 1951 a 1958. El análisis detallado de estos datos aparece en otros boletines de investigación, algunos de los cuales ya han sido publicados y otros están en preparación.